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Anadia escalerae MYERS, RIVAS & JADIN, 2009

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymAnadia escalerae MYERS, FUENMAYOR & JADIN 2009
Anadia cf. escalerae — KOK et al. 2013 
DistributionVenezuela (Bolívar), Guyana

Type locality: La Escalera region, ‘‘132 km SE Río Cuyuní (5 46 km SE km 88) on road to Santa Elena, 1430 m elevation,’’ Edo. Bolívar, Venezuela.  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: EBRG 1998, an adult female, Museo de la Estación Biológica de Rancho Grande, collected by John E. Cadle, December 30, 1980. 
DiagnosisDIAGNOSIS: Anadia escalerae most closely resembles A. pariaensis from the Península de Paria. The holotypes of pariaensis and escalerae are the only members of the genus known to have the prefrontals separated by an azygous scale (a variable character in pariaensis9 and possibly in escalerae) and these specimens also have similar color patterns (cf. figs. 9, 10). A. escalerae differs from A. pariaensis in having weakly keeled (vs. smooth) dorsal scales, in having a subocular scale protruded downward between 4th and 5th supralabials, and in differing configurations of various head scales.
 
CommentDistribution: The record from Guyana by KOK et al. 2013 is based on an enigmatic specimen collected in the Pakaraima Mountain Range and provisionally referred to Anadia escalerae. It is also the easternmost known record for an Anadia species. 
EtymologyThe specific name, a latinized noun in the genitive case, is derived from La Escalera, a well-known regional name for the steep ascent up the slope of Sierra de Lema to the northern edge of the elevated Gran Sabana, on the road from El Dorado to Santa Elena de Uairén. 
References
  • KOK, P. J. R., MEANS, D. B. & RIVAS, G. A. 2013. First record of the genus Anadia (Reptilia, Squamata, Gymnophthalmidae) from Guyana based on an enigmatic specimen from Mount Kopinang, Wokomung Massif. Boletim do Museu Paraense Emílio Goeldi 8(1): 27-39 - get paper here
  • Myers, Charles W. Rivas Fuenmayor, Gilson; Jadin, Robert C. 2009. New species of lizards from Auyantepui and La Escalera in the Venezuelan Guayana, with notes on "microteiid" hemipenes (Squamata, Gymnophthalmidae). American Museum Novitates (3660): 1-32 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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