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Andinosaura aurea (SÁNCHEZ-PACHECO, AGUIRRE-PENAFIEL & TORRES-CARVAJAL, 2012)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymRiama aurea SÁNCHEZ-PACHECO, AGUIRRE-PENAFIEL & TORRES-CARVAJAL 2012
Andinosaura aurea — SÁNCHEZ-PACHECO et al. 2017 
DistributionEcuador

Type locality: ECUADOR: El Oro: Guanazán, 2789 m elevation.  
Reproduction 
TypesHolotype: QCAZ 07886 Paratypes: El Panecillo, 2775 m (QCAZ 09649-50); Guishaguiña, Zaruma (EPNH 06196); 
DiagnosisDiagnosis (ass Riama): among the other five species of Riama currently known to occur in southern ecuador, R. au‐ rea differs from R. anatoloros in having two postparietals (three in R. anatoloros), 19-20 longitudinal dorsal scale rows in males (22-27), 34-35 transverse dorsal scale rows (36-44) and four femoral pores per hind limb in males (7-11). it differs from R. stigma‐ toral in having four femoral pores per hind limb in males (9-11 in R. stigmatoral) and eight scales between medialmost femoral pores in males (0-2). it can be distinguished from R. petrorum by the superciliary arrangement, 1+1, 1+2 or 2+2 (one, the anteriormost, in R. petrorum), by the second or second and fourth supraoculars contacting the ciliaries (second, third and fourth, or first, second and third in contact with ciliaries) and by the ventral coloration, cream with brown smears in scale centers gradually becoming more distinct ventrolaterally and on tail, forming nearly continuous longitudinal lines (venter unicolored olive or dark brown with cream along scale sutures); it further differs from R. petrorum in adult body size (maximum known svL in R. aurea is about 57 mm for males and 52 mm for females, versus 72 mm and 76 mm for males and females, respectively, of R. petrorum). from R. vespertina, R. aurea differs in having four supraoculars (three in R. vespertina) and 1+1, 1+2 or 2+2 superciliaries (2+1). from R. kiziriani it differs in having four supraoculars (three in R. kiziriani), 1+1, 1+2 or 2+2 superciliaries (2+1), four femoral pores per hind limb in males (seven) and venter cream with brown smears in scale centers gradually becoming more distinct ventrolaterally and on tail, forming nearly continuous longitudinal lines (venter dark brown to black with longitudinally arranged white stripes or spots on the scale sutures).
Riama aurea can be distinguished from the remaining ecuadorian species, as well as colombian, venezuelan, peruvian and trinidadian congeners by the number of scales between medialmost femoral pores in males (eight in R. aurea versus six or fewer in the other species) (Sanchez-Pacheco et al. 2012: 260). 
Comment 
Etymologythe specific epithet, to be treated as an indeclinable word, is an adjective derived from the Latin word aurum, meaning gold, and refers to the provenance of the species, El Oro province. 
References
  • Sánchez-Pacheco, S. , Aguirre-Peñafiel, V. , Torres-Carvajal, O. 2012. Lizards of the genus Riama (Squamata: Gymnophthalmidae): the diversity in southern Ecuador revisited. South American J. Herp. 7 (3): 259-275 - get paper here
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
 
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