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Anolis tequendama MORENO-ARIAS, MÉNDEZ-GALEANO, BELTRÁN & VARGAS-RAMÍREZ, 2023

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Tequendama anole 
SynonymAnolis tequendama MORENO-ARIAS, MÉNDEZ-GALEANO, BELTRÁN & VARGAS-RAMÍREZ 2023: 179
Anolis heterodermus — CASTAÑEDA & DE QUEIROZ 2013 
DistributionColombia (Cundinamarca)

Type locality: the Vereda Sabaneta, San Francisco municipality, Cundinamarca department, Colombia (4.891173°N –74.289925°W, 2850 m),  
Reproduction 
TypesHolotype. ICN 4548. Adult male with everted hemipenis, dewlap present, enlarged postanal scales, collected by Henry Zuñiga and Jane R. Rodríguez in 1981.

Paratypes. ICN 4546 and 4547 adult male and female, respectively from the same locality data of the holotype, collected by Jose Vicente Rueda in 1981. ICN 5734 adult female from Cueva Grande hill From the Vereda Sabaneta, San Francisco municipality, Cundinamarca department, Colombia, collected by Henry Zúñiga in 1981. ICN-1499 adult male from the Vereda de Fute, Las Mercedes, Bojacá municipality, Cundinamarca department, Colombia (4.616388°N –74.281944°W, 2600 m), collected by J. Hernandez-Camacho in 1962. ICN 5765 (adult female) from Laguna de Pedro Palo, Tena municipality, Cundinamarca department, Colombia (4.683888°N –74.387222°W, 2050 m), collected by Oscar Pinto in 1981. ICN 9688 (adult female) from Barro Colorado, La Agüadita, Fusagasugá municipality, Cundinamarca department, Colombia (4.388888°N –74.334444°W, 1800 m), collected by D. Rivera in 1990. ICN 10609 (adult male) from Granjas del Padre Luna, Alban municipality, Cundinamarca department, Colombia (4.899133°N –74.424122°W, 2090 m), collected by John D. Lynch in 1985. 
DiagnosisDiagnosis: Anolis tequendama sp. nov. shares short limbs, a large casqued head, a prehensile tail without caudal autotomy, and lamellar subdigital scales of all digits extending from the most proximal phalanges with the other species of the Phenacosaurus clade. Anolis tequendama sp. nov. differs from A. proboscis, A. orcesi, A. euskalerriari and A. nicefori, by its strongly heterogeneous dorsal scalation. Anolis tequendama sp. nov. differs from A. nicefori by the presence of granular scales surrounding dorsal heterogeneous flat scales. A. tequendama sp. nov. differs from A. heterodermus, A. richteri, A. tetarii, A. inderenae, A. vanzolinii and A. quimbaya sp. nov. by a male dewlap colour uniformly brown or brown bluish. Anolis tequendama sp. nov. further differs from A. inderenae, A. heterodermus, A. richteri and A. tetarii by the presence of a discontinuous nuchal crest, from A. heterodermus and A. inderenae by the presence of a continuous row of enlarged sublabials that do not reach the mouth commissure; from A. heterodermus by a V-shaped crown; and from A. vanzolinii by < 24 expanded lamellae under second and third phalange of fourth toe and < 35 lamellae in total fourth toe.


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Comment 
EtymologyNamed after the Tequendama Falls, a popular waterfall near Bogotá that is part of the Bogotá River and has cosmological importance for the Muisca indigenous people; inhabitants of the Bogotá Plateau. In the Muisca language (called “Muysccubun”), Tequendama means “he who precipitates downward”, referring to the waterfall. 
References
  • Castañeda, Rosario María Del and KEVIN DE QUEIROZ 2013. Phylogeny of the Dactyloa Clade of Anolis Lizards: New Insights from Combining Morphological and Molecular Data. Bull. Mus. Comp. Zool. Harvard 160 (7): 345-398. - get paper here
  • Moreno-Arias, R. A., Méndez-Galeano, M. A., Beltrán, I., & Vargas-Ramírez, M. 2023. Revealing anole diversity in the highlands of the Northern Andes: New and resurrected species of the Anolis heterodermus species group. Vertebrate Zoology, 73, 161-188 - get paper here
 
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