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Anolis scypheus COPE, 1864

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Yellow-tongued Anole
Portuguese: Papa-Vento 
SynonymAnolis scypheus COPE 1864: 172
Anolis chrysolepis — GUICHENOT 1855: 15
Anolis scypheus — BOULENGER 1885: 90
Anolis incompertus incompertus BARBOUR 1932: 99
Anolis incompertus nicefori BARBOUR 1932: 100
Anolis scypheus — DONOSO-BARROS 1968
Anolis chrysolepis scypheus — VANZOLINI & WILLIAMS 1970
Anolis chrysolepis scypheus — DUELLMAN 1978: 198
Norops nitens scypheus — SAVAGE & GUYER 1991: 366
Anolis nitens scypheus — AVILA-PIRES 1995
Anolis nitens scypheus — GRAHAM 2000
Anolis nitens scypheus — WHITHWORTH & BEIRNE 2011
Anolis scypheus — D’ANGIOLELLA et al. 2011
Norops scypheus — NICHOLSON et al. 2012
Norops scypheus — NICHOLSON et al. 2018 
DistributionAmazonian areas of Colombia, Ecuador, Peru, Brazil (Amazonas)

Type locality: ‘‘Caracas’’ according to Boulenger, 1885, but considered an error by Vanzolini and Williams, 1970:85) or doubtful (HOOGMOED, pers. comm.); None given according to PETERS & DONOSO-BARROS 1970.  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.8.55
Holotype: MCZ 32309 [incompertus]
Holotype: MCZ 32322 [nicefori] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (551 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: Williams 1970 synonymized A. incompertus without mentioning A. (i.) nicefori, hence this synonymy is preliminary.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
EtymologyNamed after Greek scyphos (σκῦφος), skull. (COPE 1864, Esteban Lavilla, pers. comm., April 2024) 
References
  • Avila-Pires, T.C.S. 1995. Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1-706 - get paper here
  • Barbour, Thomas 1932. New Anoles. Proc. New England Zool. Club 12: 97-102
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Cope, E.D. 1864. Contributions to the herpetology of tropical America. Proc. Acad. Nat. Sci. Philadelphia 16: 166-181. - get paper here
  • Crnobrna B, Santa-Cruz Farfan R, Gallegos C, López-Rojas JJ, Llanqui IB, Pisco GP, Arbaiza AK 2023. Herpetological records from the Abujao basin, central Peruvian Amazon. Check List 19(4): 433-465 - get paper here
  • D'angiolella, Annelise B.; Tony Gamble, Teresa C. S. Avila-Pires, Guarino R. Colli, Brice P. Noonan, and Laurie J. Vitt 2011. Anolis chrysolepis Duméril and Bibron, 1837 (Squamata: Iguanidae), Revisited: Molecular Phylogeny and Taxonomy of the Anolis chrysolepis Species Group. Bull. Mus. Comp. Zool. Harvard 160 (2): 35-63. - get paper here
  • Dixon, J. R.; Soini, P. 1975. The reptiles of the upper Amazon basin, Iquitos region, Peru, Part I. Lizards and Amphisbaenians. Milwaukee Public Museum Contributions in Biology and Geology 4: 1-58 [1986?] - get paper here
  • Donoso-Barros, R. 1968. The lizards of Venezuela (checklist and key). Carib. J. Sci. 8 (3-4): 105-122
  • Duellman, W. E. 1978. The biology of an equatorial herpetofauna in Amazonian Ecuador. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 65: 1-352 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Graham, D.L. 2000. Annotated list of reptiles and amphibians recorded from the Rio Orosa and surrounding areas, Department of Loreto, Peru. http://www.projectamazonas.com/doc/herps.pdf
  • Guichenot, A. 1855. Animaux nouveaux ou rares recueillis pendant l’Expédition dans les parties centrales de l’Amérique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; Exécutée par ordre du Gouvernement Francais pendant les années 1843 a 1847, sous la direction du P. Bertrand, Paris. - get paper here
  • Medina-Rangel, Guido Fabian; Miguel Á. Méndez-Galeano &<br>Martha Lucía Calderón-Espinosa 2019. Herpetofauna of San José del Guaviare, Guaviare, Colombia Biota Colombiana 20 (1): 75-90 - get paper here
  • Muñoz-Saba, Yaneth; Nathaly Calvo-Roa, Paul Andrés Gómez-Sandoval, Diego Casallas-Pabón John Douglas Lynch, Lucas S. Barrientos, Diego A. Gómez-Sánchez. 2019. Guía de campo de los mamíferos, anfibios y reptiles de Santa María (Boyacá, Colombia). Serie Guías de Campo del Instituto de Ciencias Naturales. N° 23. Bogotá D.C. Universidad Nacional de Colombia. Bogotá. 317 p - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983 (1): 001–110 - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • Rodríguez-Rodríguez, J. D., & Calderón-Espinosa, M. L. 2024. Sexual dimorphism and female ecomorphology in Anolis (Squamata: Anolidae): knowledge of female morphology increases the understanding of Anolis diversification. Biological Journal of the Linnean Society, blae019 - get paper here
  • Savage,J.M. & Guyer,C. 1991. Nomenclatural notes on Anoles (Sauria: Polychridae): stability over priority. Journal of Herpetology 25 (3): 365-366 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Vanzolini,P.E. & Williams,E.E. 1970. South American anoles: the geographic differentiation and evolution of the Anolis chrysolepis species group (Sauria, Iguanidae). Arquivos de Zoologia (São Paulo) 19: 1-298 - get paper here
  • Whithworth, A. & Beirne, C. 2011. Reptiles of the Yachana Reserve. Global Vision International, 130 pp. - get paper here
 
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