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Anolis lyra POE, VELASCO, MIYATA & WILLIAMS, 2009

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymAnolis lyra POE, VELASCO, MIYATA & WILLIAMS 2009
Norops lyra — NICHOLSON et al. 2012
Norops lyra — NICHOLSON et al. 2018 
DistributionC Colombia (Pacific lowlands), NW Ecuador

Type locality: Ecuador, Pichincha, Finca Victoria, 37 km SE of Santo Domingo.  
Reproductionoviparous 
TypesHolotype: MCZ 80955, adult female, 16 July 1964, F. Vuilleumier. 
DiagnosisDiagnosis: Anolis lyra differs from all other South American Anolis except A. vittigerus in possessing a small red dewlap with a dark central spot in males and a lyre-shaped symbol in the occipital region (Fig. 8). Anolis lyra differs from A. vittigerus in possessing smaller middorsal scales (mean number of longitudinal scales in 5% SVL for A. lyra 5 8.5, A. vittigerus 5 5.8; P 5 0.006, MannWhitney U test), more scales between supraorbital semicircles (mean 5 2.3 in A. lyra, 0.6 in A. vittigerus; P 5 0.001, Mann-Whitney U test), and more scales between interparietal and supraorbital semicircles (x¯ 5 4.1 in A. lyra, 2.4 in A. vittigerus; P 5 0.001, MannWhitney U test; Fig. 10). Also, A. lyra tends to display much weaker lateral striping (i.e., broken lines only extending to the axillae versus broad stripes to groin in A. vittigerus) and a more prominent lyre-shaped posterior head marking (Fig. 8; absent to weak in A. vittigerus). In western Colombia, the only area in which both species can be expected to occur, A. lyra and A. vittigerus may be most easily separated by the condition of the supraorbital semicircles—in broad contact in A. vittigerus; usually separated by 2–3 scales in A. lyra.
 
CommentSpecies group: Norops auratus Species Group (fide Nicholson et al. 2012) 
EtymologyThis species is named for the lyre-shaped marking on the back of its head. 
References
  • Arteaga AF, Bustamante-Enríquez LM and Guayasamin JM 2013. The Amphibians and Reptiles of Mindo. http://www.tropicalherping.com - get paper here
  • Carvajal-Cogollo, J.E.; L.E. Rojas-Murcia. & G. Cárdenas-Arévalo 2020. Reptiles del Caribe colombiano/ Reptiles of the Colombian Caribbean. Tunja: Editorial UPTC, 268 pp. - get paper here
  • Chun, Wesley 2010. Miscellaneous notes on some rare and unusual anoles. Anolis Newsletter VI: 14-22 - get paper here
  • Cortés Gómez, Angela María; Anyelet Valencia Aguilar, Diana Marcela Torres Domínguez, Lina María García Calderón, Diego Fernando Villaquirán Martínez, Andrea del Pilar Cáceres Franco, Fernando Castro Herrera 2010. Guía de los anfibios y reptiles. Área en conservación de la microcuenca Quebrada Pericos. [Valle de Cauca, Colombia]. Corporación Autónoma Regional del Valle del Cauca, Santiago de Cali: CVC, 37 pp. - get paper here
  • Cruz-García K, Cisneros-Heredia DF, Sánchez-Nivicela JC, Amador L 2020. New records of four reptile species (Lepidosauria, Squamata) from the province of Guayas, southwestern Ecuador. Check List 16(3): 765-772 - get paper here
  • HARRISON, ALEXIS; STEVEN POE 2012. Evolution of an ornament, the dewlap, in females of the lizard genus Anolis. Biological Journal of the Linnean Society 106: 191–201 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Pinto-Erazo, M.A., Calderón-Espinosa, M.L., Medina-Rangel, G.F. & Méndez-Galeano, M.A. 2020. Herpetofauna from two municipalities of southwestern Colombia. Biota Colombiana 21(1): 41-57. DOI: 10.21068/c2020.v21n01a04 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Poe, S.; Velasco, J.; Miyata, K. & Williams, E.E. 2009. Descriptions of two nomen nudum species of Anolis lizard from Northwestern South America. Breviora (516): 1-16 - get paper here
  • Rengifo M, Jhon Tailor; Fernando C. Herrera, Francisco J. Purroy 2014. Diversidad de una comunidad de Anolis (Iguania: Dactyloidae) en la selva pluvial central, departamento del Chocó, Colombia. Basic and Applied Herpetology 28: 51-63 - get paper here
  • Rengifo M., J. T., Castro Herrera, F. & Purroy Iraizos, F. J. 2015. Uso de hábitat y relaciones ecomorfológicas de un ensamble de Anolis (Lacertilia: Dactyloidae) en la región natural Chocoana, Colombia. Acta Zoológica Mexicana (n.s.) 31 (2): 159-172 - get paper here
  • Thomas, O., Preston, S., Cole, T., McDonald, R. V., Castellano, I., & Neira-Salamea, K. 2023. Herps at height: Records of seven species utilizing the upper canopy in northwestern Ecuador. Reptiles & Amphibians, 30(1), e19692-e19692
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Valencia-Zuleta A, Jaramillo-Martínez AF, Echeverry-Bocanegra A, Viáfara-Vega R, Hernández-Córdoba O, Cardona-Botero VE, Gutiérrez-Zúñiga J, Castro-Herrera F. 2014. Conservation status of the herpetofauna, protected areas, and current problems in Valle del Cauca, Colombia. Amphibian & Reptile Conservation 8 (2): 1–18 (e87) - get paper here
  • Vargas-S., F. & Bolanos-L., M.E. 1999. Presencia de reptiles en la región de Anchicayá, pacifico colombiano, a través de un gradiente de deforestación. Caldasia 21 (2): 235-238 - get paper here
  • Yánez-Muñoz, M. H.; J. C. Sánchez L., K. López, E. Rea S., P. A. Meza-Ramos, L. A. Oyagata C., P. Guerrero 2014. Ampliaciones del rango de distribución de algunas especies de anfibios y reptiles en el suroccidente de Ecuador. Avances en Ciencias e Ingenierías 6 (1): B2-B5 - get paper here
 
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