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Anolis bremeri BARBOUR, 1914

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
SubspeciesAnolis bremeri bremeri BARBOUR 1914
Anolis bremeri insulaepinorum GARRIDO 1972 
Common NamesE: Cuban Variegated Anole, Herradura Anole 
SynonymAnolis bremeri BARBOUR 1914: 288
Norops bremeri — SCHWARTZ & HENDERSON 1988
Anolis bremeri — SCHWARTZ & HENDERSON 1991: 230
Anolis bremeri — RODRÍGUEZ SCHETTINO 1999: 328
Norops bremeri — NICHOLSON 2002
Norops bremeri — NICHOLSON et al. 2012
Norops bremeri — NICHOLSON et al. 2018

Anolis bremeri insulaepinorum GARRIDO 1972
Anolis bremeri insulaepinorum GARRIDO 1972
Norops bremeri insulaepinorum — SCHWARTZ & HENDERSON 1988
Anolis bremeri insulaepinorum — POWELL & HENDERSON 2012: 124 
DistributionCuba (Pinar del Rio), Isla de la Juventud

bremeri: Cuba, Pinar del Rio Prov.; Type locality: Herradura, Pinar del Rio Province, Cuba.

insulaepinorum: Isla de la Juventud, north of Ciénaga de Lanier.  
TypesHolotype: MCZ 7889
Holotype: CZACC = IZ 1626 [insulaepinorum] 
DiagnosisDESCRIPTION: Size moderate (SVL in males to 72 mm, in females to 52 mm); 5
loreal rows; scales between supraorbitals 0-2 (strongly modally 1); 2-4 scales between interparietal and supraorbital semicircles; 5 postrostrals; 1-8 postmentals; dorsals smooth, flat, nonimbricate, a middorsal zone about 6 scales wide slightly enlarged, imbricate; lateral scales small, granular but keeled; ventrals keeled, imbricate, acuminate, in longitudinal and diagonal rows; supradigital scales multicarinate; tail laterally compressed, verticillate, scales in 7 vertical rows/verticil, upper edge feebly serrate with 4 small spines/verticil; females with snout distinctly recurved or upturned. Dorsum grayish brown to brown with sides yellow, with many cream-colored bands, variegated with white dots and lines, and about 4 parallel lines on sides of venter; venter sandy; limbs with cream bands; dewlap very large, ochraceous or mustard to mahogany, interior scales black, those near throat white; iris brown; females with pale dorsal rhombs (Schwartz & Henderson 1991: 230). 
CommentIllustrations: Barbour and Ramsden, 1919; Peters and Schubert, 1968; Garrido, 1972.

Subspecies: Anolis bremeri insulaepinorum GARRIDO, 1972 is not listed by NICHOLSON et al. 2012, 2018. It may be a synonym of bremeri or some other species.

Species group: Norops sagrei species group (fide NICHOLSON et al. 2012). 
EtymologyNamed after Dr. John Lewis Bremer (1874-1959) who worked at the Harvard Medical School as Professor of Anatomy (1931-). Barbour wrote that Bremer "has often most kindly aided me while upon collecting trips." 
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Barbour, Thomas 1914. A Contribution to the Zoögeography of the West Indies, with Especial Reference to Amphibians and Reptiles. Memoirs of the Museum of Comparative Zoölogy 44 (2): 205-359 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Garrido, O.H. 1972. Anolis bremeri Barbour (Lacertilia: Iguanidae) en el occidente de Cuba e Isla de Pinos. Carib. J. Sci. 12 (1-2): 59-77.
  • Nicholson, K.E. 2002. Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs 16: 93-120 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Peters, G.,and Schubert, R. 1968. Theoretische und praktische Aspekte einer biologischen Expedition durch Kuba. Biol. Schule 8/9:337-46.
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
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