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Anolis kemptoni DUNN, 1940

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Kempton's Anole 
SynonymAnolis kemptoni DUNN 1940
Anolis kemptoni — PETERS & DONOSO-BARROS 1970: 57
Anolis kemptoni — WILLIAMS et al. 1995
Norops pandoensis SAVAGE & GUYER 1998
Norops pandoensis — KÖHLER 2000: 63
Norops kemptoni — KÖHLER 2000: 62
Norops kemptoni — NICHOLSON 2002
Norops pandoensis — NICHOLSON 2002
Norops pandoensis — SVAGE 2002: 473
Norops pandoensis — NICHOLSON et al. 2005
Norops kemptoni — NICHOLSON et al. 2012
Norops kemptoni — NICHOLSON et al. 2018 
DistributionPanama (Chiriquí), elevation 1600-1900 m

Type locality: Finca Lérida, Chiriquí, Panama, 5300 ft. elevation.

pandoensis: Costa Rica (Cordillera de Talamanca), Panama; Type locality: Costa Rica: Puntarenas Province: Cantón Coto Brus: Zona Protectora Las Tablas: La Changuinolita, elevation 1950 m.  
TypesHolotype: ANSP 21708
Holotype: Costa Rican Expeditions (CRE) collection 5290, an adult male; collected by Karen R. Lips, September 21, 1991 [pandoensis] 
DiagnosisDiagnosis (pandoensis): A small, yellowish-tan, short-legged member of the Norops fuscoauratus species group characterized by having enlarged postcloacals and a large, bicolored pink and orange dewlap in males and a small, plain dewlap not contrasting in color with the rest of the throat in females. [from SAVAGE & GUYER 1998].

Comparisons: N. pandoensis is a member of the fuscoauratus species group. Lizards of this group are small to moderate in size, have small head scales, small dorsal scales, mostly smooth ventrals, long tails that are round to subcylindrical in cross-section. Members of this group usually have an inscriptional rib formula of 3: 1 (Etheridge, 1965) as is the case with N. pandoensis.
Among Central American members of the fuscoauratus group the new species is unlikely to be confused with Norops limifrons (including N. biscutiger) or Norops polylepis both of which have long legs, i.e., the tip of the 4th toe reaches beyond the orbit when the leg is adpressed along the side of the body. In addition, the dewlap in male N. limifrons is white with a basal yellow to orange spot and is uniform orange in male N. polylepis.
Among short-legged members of the fuscoauratus group, N. pandoensis can be distinguished immediately from the lowland Norops carpenteri which is green, smaller (standard length 35-45 mm) and with a uniform orange dewlap color in males. Comparisons with the four short-legged species most similar to N. pandoensis in general appearance are summarized in Table 1. All are upland forms and one or more of them may prove to occur sympatrically with N. pandoensis. [from SAVAGE & GUYER 1998]. 
CommentSynonymy: Norops pandoensis has been synonymized with N. kemptoni by HULEBAK & POE 2006.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
EtymologyNamed after Dr. Kempton Potter Aiken Taylor, an American physician who was born Kempton Potter Aiken but took the surname Taylor after adoption. He worked in Guatemala (1928) and Panama, where he met Dunn, an old friend. 
  • Batista, A., Hertz, A., Ponce, M. and Lotzkat, S. 2020. Notes on amphibians and reptiles from western Panama Notes on amphibians and reptiles from western Panama. Herpetology Notes 13: 219-229. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Dunn,E.R. 1940. New and noteworthy herpetological material from Panamá. Proc. Acad. Nat. Sci. Philadelphia. 92: 105-122. - get paper here
  • Echelle,A.A.; Echelle,A.F. & FITCH,H.S. 1971. A new anole from Costa Rica. Herpetologica 27: 354-362 - get paper here
  • Hulebak, E. & Poe, S. 2006. Anolis (Norops) pandoensis Savage and Guyer 1998 Is a Junior Synonym of Anolis kemptoni Dunn 1940. Carib. J. Sci. 42 (2): 268-270
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Kohler, G. 2011. A new species of anole related to Anolis altae from Volcán Tenorio, Costa Rica (Reptilia, Squamata, Polychrotidae). Zootaxa 3120: 29-42 - get paper here
  • Köhler, Gunther 2009. New Species of Anolis Formerly Referred to as Anolis altae from Monteverde, Costa Rica (Squamata: Polychrotidae). Journal of Herpetology 43 (1):11-20 - get paper here
  • Köhler, Gunther, Joseph Vargas, Johannes J. Köhler and Milan Vesel´y. 2013. Noteworthy distributional records of amphibians and reptiles from Costa Rica. Herpetological Review 44 (2): 280-283 - get paper here
  • Mata-Silva, V, DeSantis DL, García-Padilla E, Johnson JD, Wilson LD. 2019. The endemic herpetofauna of Central America: a casualty of anthropocentrism. Amphibian & Reptile Conservation 13 (1) [General Section]: 1–64 (e168) - get paper here
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos 2005. Mainland colonization by island lizards. Journal of Biogeography 32: 929–938 - get paper here
  • Nicholson, K.E. 2002. Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs 16: 93-120 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Ponce, M. & Köhler, G. 2008. Morphological variation in anoles related to Anolis kemptoni in Panama. Salamandra 44 (2): 65-83 - get paper here
  • Savage, J. M. & C. GUYER 1998. A new species of anole lizard, genus Norops (Squamata: Polychrotidae), from the Cordillera de Talamanca, Costa Rica. Revista de Biologia Tropical 46 (3): 805-809. - get paper here
  • Schlüter, U. 2001. Neuerungen in der Systematik der mittelamerikanischen Saumfinger der Gattung Norops seit 1996. Iguana Rundschreiben 14 (2): 19-28
  • Williams, E.E.; Rand, H.; Rand, A.S. & O’Hara, R.J. 1995. A computer approach to the comparison and identification of species in difficult taxonomic groups. Breviora (502): 1-47 - get paper here
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