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Anolis krugi PETERS, 1877

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Common NamesOlive bush anole, Krug's Anole, Orange Dewlap Anole

Spanish: Lagartijo de las yerbas de la montaña 
SynonymAnolis Krugi PETERS 1877: 707
Anolis krugi — BOULENGER 1885: 37
Anolis krugi — FOWLER 1918
Anolis krugi — GRANT 1932
Anolis krugi — SCHWARTZ & HENDERSON 1991: 285
Anolis krugi — BRANDLEY & DE QUEIROZ 2004
Ctenonotus krugi — NICHOLSON et al. 2012
Ctenonotus krugi — NICHOLSON et al. 2018 
DistributionPuerto Rico

Type locality: “ohne genauere Angabe des Fundorts” [i.e. not given].  
TypesHolotype: ZMB 8965 
DiagnosisDESCRIPTION: Size small (SVL in males to 45 mm, in females to 36 mm); habitus
rather attenuate; 5-7 rows of loreals; scales between supraorbitals 1-2; scales between interparietal and supraorbital semicircles 2-4; 5 postrostrals; 4 postmentals; suboculars in contact with supralabials; head scales keeled or wrinkled; about 6 middorsal rows enlarged, keeled, imbricate, remainder of back and sides with minute granules; ventrals large, imbricate, strongly keeled, keels forming continuous ridges, scales on throat smaller and narrower; supradigital scales multicarinate; digits (especially toes) long and slender; tail very long, more than 2 x SVL, moderately compressed, with large, subequal keeled scales, nonverticillate, middorsals forming a feebly serrate ridge; dewlap edge thickened. Dorsum (male) bright, yellowish olive-green, almost yellow, or almost black; sides of back and sides of body with small black spots, but none on area of enlarged middorsals; a broad, canary-yellow to cream or white stripe from below eye to groin, bright on middle of flanks; below stripe, lower sides olivaceous, with minute black flecks; hindlimbs suffused with ferruginous; tail crossbarred with black and buffy; a black postocular spot; venter buffy to pinkish; dewlap yellowish, orange, or peach toward edge; eyelids edged with whitish; iris very dark brown (nearly black) with faint, silvery pupillary ring; female dark olive-brown, paler below straw-colored lateral stripe; venter pale yellowish; median dorsal zone in juveniles strongly
scalloped, white (Schwartz & Henderson 1991: 285). 
CommentFor illustrations see Stejneger, 1904; Schmidt, 1928; Rivero, 1978; Schwartz and Henderson, 1985.
Ecomorph class: grass-bush.

Species group: Ctenonotus cristatellus species group (fide NICHOLSON et al. 2012). 
EtymologyNamed after Carl (Karl) Wilhelm Leopold Krug (1833-1898), a German businessman who went to work in Puerto Rico where he became the German Vice Consul. He collected the holotype of this species together with Gundlach. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Brandley, M.C. & De Queiroz, K. 2004. Phylogeny, ecomorphological evolution, and historical biogeography of the Anolis cristatellus series. Herpetological Monographs 18: 90-126 - get paper here
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Fowler, H. W. 1918. Some amphibians and reptiles from Porto Rico and the Virgin Islands. Publ. Carnegie Inst. Washington, 252: 1-16.
  • Grant, C. 1932. Herpetological notes from the Puerto Rico area. J. Dept. Agric. Puerto Rico 16 (1): 161-165
  • Horstink, Yann 2004. Anolis krugi - Een kleine grasanolis van Puerto Rico. Lacerta 62 (2): 60-65 - get paper here
  • Medina, Iliana; Jonathan B. Losos andD. Luke Mahler 2016. Evolution of dorsal pattern variation in Greater Antillean Anolis lizards. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12881 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Peters, W. 1877. Über eine von Hrn. Viceconsul L. Krug und Dr. J. Gundlach auf der Insel Puertorico gemachte Sammlung von Säugethieren und Amphibien, so wie über die Entwickelung eines Batrachiers, Hylodes martinicensis Dum. Bibr., ohne Metamorphose. Monatsber. Königl. Preuss. Akad. Wiss. Berlin, 1876: 703-714. - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rios-López, N. & Aide, T.M. 2007. Herpetofaunal dynomics during secondary succession. Herpetologica 63 (1): 35-50 - get paper here
  • Rivero, J.A. 1978. Los anfibios y reptiles de Puerto Rico. M. Pareja Montana, 16, Barcelona, Espafia: x + 152 + 148pp.
  • Schmidt, K.P. 1928. Scientific Survey of Porto Rico and the Virgin Islands: Amphibians and land reptiles of Porto Rico, with a list of those reported from the Virgin Islands. New York Academy of Sciences 10 (1):160 pp.
  • Schoener, Thomas W.;Schoener, Amy 1971. Structural habitats of West Indian Anolis lizards II. Puerto Rican uplands. Breviora (375): 1-39 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Henderson, R.W. 1985. A guide to the identification of the amphibians and reptiles of the West Indies exclusive of Hispaniola. Milwaukee Public Mus., 165 pp.
  • Stejneger, L. 1904. The herpetology of Porto Rico. Rept. United States Natl. Mus. 1902: 549-724.
  • Thonis, Anna E.; Bradford C. Lister 2019. Predicting Climate-Induced Distributional Shifts for Puerto Rican Anoles. Copeia 107 (2): - get paper here
  • Werning, H. 2012. Zwischen Anolis und Cycluren: Unterwegs auf Puerto Rico. Reptilia (Münster) 17 (95): 100-109 - get paper here
  • Yuan, M. L., M. H. Wake, and I. J. Wang. 2019. Phenotypic integration between claw and toepad traits promotes microhabitat specialization in the Anolis adaptive radiation. Evolution 73: 231–244 - get paper here
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