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Anolis loysiana DUMÉRIL & BIBRON, 1837

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Spiny Anole, Peach Anole 
SynonymAnolis loysiana DUMÉRIL & BIBRON 1837: 100
Heteroderma loysianum — FITZINGER 1843: 68
Acantholis loysiana — GRAY 1845: 206
Anolis loysiana — BOULENGER 1885: 42
Anolis loysiana — WILLIAMS 1976: 14
Anolis loysianus — SCHWARTZ & HENDERSON 1988: 82
Anolis loysianus — SCHWARTZ & HENDERSON 1991: 291
Anolis loysiana — RODRÍGUEZ SCHETTINO 1999: 243
Anolis loysiana — LOSOS et al. 2003
Anolis loysianus — NICHOLSON et al. 2012
Anolis loysianus — NICHOLSON et al. 2018 
DistributionCuba

Type locality: Cuba.  
Reproductionoviparous 
TypesHolotype: MNHN-RA 2465 (by monotypy); note that MNHN-RA catalogues indicated "louisiana" and not loysiana 
DiagnosisDESCRIPTION: Size small (SVL in males to 40 mm, in females to 37 mm); 4 rows of loreals; 0 scales between supraorbitals; 1 scale between interparietal and supraorbital semicircles; 5 postrostrals; 6 postmentals; suboculars in contact with supralabials; supradigital scales smooth; dorsals small, flat, smooth, 2 middorsal rows slightly enlarged; dorsum and limbs with scattered, large, spine-like scales; head scales smooth; sides with small flat scales and with projecting groups of scales, apical ones elongate, spine-like; ventrals small, flat, slightly imbricating; tail cylindrical, about equal to SVL, with irregular median dorsal series of enlarged imbricating scales, many scattered, spine-like scales, nonverticillate. Dorsum very light gray with fine narrow crossbands, anastomosing lines, and spots dark gray or mauve; venter white; dewlap tan to pinkish tan, pale orange-red, or pale yellow (with scattered orange spots) (Schwartz & Henderson 1991: 291). 
CommentFor illustrations see Barbour and Ramsden, 1919, Flaschendräger 2002.

Species group: Anolis loysianus species group (fide NICHOLSON et al. 2012). 
References
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fitzinger, L. 1843. Systema Reptilium, fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp. - get paper here
  • Flaschendräger, A. 2002. Beobachtungen an Anolis-Arten in der Cordillera de Guaniguanico, Provinz Pinar del Río, West-Kuba. Herpetofauna 24 (138): 5-18 - get paper here
  • Fläschendräger, Axel 2017. Beobachtungen an einigen Anolis-Arten im westlichen Kuba. Terraria-Elaphe 2017 (5): 52-63 - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Losos, Jonathan B., Manuel Leal, Richard E. Glor, Kevin de Queiroz, Paul E. Hertz, Lourdes Rodríguez Schettino, Ada Chamizo Lara, Todd R. Jackman & Allan Larson 2003. Niche lability in the evolution of a Caribbean lizard community. Nature 424: 542-545 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Ruibal, R. 1964. An annotated checklist and key to the anoline lizards of Cuba. Bull. Mus. comp. Zool. Harvard 130 (8): 475-520. - get paper here
  • Schwartz, A. & Henderson, R.W. 1988. West Indian Amphibian and Reptiles: A Checklist. Milwaukee Publ. Mus. Contr. Biol. Geol. No. 74: 264 pp.
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • WERNING, Heiko 2012. Im Bad mit Anolis – ein Besuch bei Uwe Bartelt. Terraria-Elaphe 2012 (6): 88-92 - get paper here
  • Williams, E. E.; Rand, A. S. 1977. Species recognition, dewlap function and faunal size. American Zoologist 17: 261-270 - get paper here
  • Williams,E.E. 1976. West Indian Anoles: a taxonomic and evolutionary summary. 1. Introduction and a species list. Breviora (440): 1-21 - get paper here
  • Yuan, M. L., M. H. Wake, and I. J. Wang. 2019. Phenotypic integration between claw and toepad traits promotes microhabitat specialization in the Anolis adaptive radiation. Evolution 73: 231–244 - get paper here
 
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