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Anolis planiceps TROSCHEL, 1848

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Golden-scaled Anole, Orange-fanned leaf-litter anole, Goldenscale Anole
S: Anolis común, anolis camaleoncito
Portuguese: Papa-Vento 
SynonymAnolis planiceps TROSCHEL 1848: 649
Anolis eewi ROZE 1958: 311
Anolis eewi — PETERS et al. 1970: 52
Anolis chrysolepis planiceps — VANZOLINI & WILLIAMS 1970: 85
Anolis nitens nitens — AVILA-PIRES 1995
Anolis chrysolepis eewi — WILLIAMS et al. 1996: 10
Anolis chrysolepis planiceps — MYERS & DONNELLY 1997
Anolis eewi — GORZULA & SEÑARIS 1999
Anolis chrysolepis planiceps — MYERS & DONNELLY 2008
Anolis planiceps — D’ANGIOLELLA et al. 2011
Norops eewi — NICHOLSON et al. 2012
Norops planiceps — NICHOLSON et al. 2018 
DistributionE Venezuela (Bolivar, Amazonas), Guyana, Brazil (Roraima, Amazonas), Trinidad.

Type locality: Caracas, Venezuela

(Sub-) species no longer recognized:

eewi: Venezuela (Bolivar); Type locality: Cumbe de Torono Tepui, Chimantá Tepui, Estado Bolívar, Venezuela, 2100 m elevation.  
TypesHolotype: ZMB 529
Holotype: FMNH 74040 [eewi] 
DiagnosisDescription. Maximum SVL 76 mm. Double row of enlarged vertebral scales extending from nape to base of tail; few to several rows of weakly keeled scales, increasing in number caudally, forming a gradual transition between double row of enlarged scales and granules on flanks. Scales of upper arms markedly larger than vertebral scales. Supraorbital semicircles with enlarged scales, forming pronounced ridge in some specimens. Supraocular region with distinct group of enlarged, weakly keeled, scales surrounded by smaller scales. Interparietal distinctly larger than adjacent scales [from D’ANGIOLELLA et al. 2011]. 
CommentNot mentioned at all by NICHOLSON et al. 2012.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012)

Synonymy after Peters et al. 1970, VANZOLINI & WILLIAMS 1970, and Poe et al. 2017. Gorzula and Señaris (1999) noted that Anolis eewi is distinguished from A. chrysolepis by shorter legs but this character is somewhat variable, which is why Myers and Donnelly 2008 supported to synonymize eewi and planiceps.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
EtymologyNamed after Latin “planus” = flat and “-ceps” head.

A. eewi was named after Ernest E. Williams, who was often known by his initials, EEW. 
  • Auguste, Renoir J. 2019. Herpetofaunal checklist for six pilot protected areas in Trinidad and Tobago. Herpetology Notes 12: 577-585 - get paper here
  • Avila-Pires, T.C.S. 1995. Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1-706 - get paper here
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • D'angiolella, Annelise B.; Tony Gamble, Teresa C. S. Avila-Pires, Guarino R. Colli, Brice P. Noonan, and Laurie J. Vitt 2011. Anolis chrysolepis Duméril and Bibron, 1837 (Squamata: Iguanidae), Revisited: Molecular Phylogeny and Taxonomy of the Anolis chrysolepis Species Group. Bull. Mus. Comp. Zool. Harvard 160 (2): 35-63. - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Gorzula, Stefan & Senaris, J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Moraes LJCL, Almeida AP, Fraga R, Rojas RR, Pirani RM, Silva AAA, de Carvalho VT, Gordo M, Werneck FP 2017. Integrative overview of the herpetofauna from Serra da Mocidade, a granitic mountain range in Northern Brazil . ZooKeys 715: 103-159 - get paper here
  • Myers, Charles W.;Donnelly, Maureen A. 1997. A Tepui Herpetofauna on a Granitic Mountain (Tamacuari) in the Borderland Between Venezuela and Brazil: Report from the Phipps Tapirapecó Expedition. American Museum Novitates (3213): 1-71 - get paper here
  • Myers, Charles W.;Donnelly, Maureen A. 2008. The Summit Herpetofauna Of Auyantepui, Venezuela: Report From The Robert G. Goelet American Museum-Terramar Expedition. Bulletin of the American Museum of Natural History 308: 1-147 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Oliveira, D.P.; S.M. Souza; L. Frazão; A.P. Almeida; T. Hrbek 2014. Lizards from central Jatapú River, Amazonas, Brazil. Check List 10 (1): 46-53 - get paper here
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Poe, Steven; Adrián Nieto-montes de Oca, Omar Torres-carvajal, Kevin De Queiroz, Julián A. Velasco, Brad Truett, Levi N. Gray, Mason J. Ryan, Gunther Köhler, Fernando Ayala-varela, Ian Latella 2017. A Phylogenetic, Biogeographic, and Taxonomic study of all Extant Species of Anolis (Squamata; Iguanidae). Syst Biol 66: 663–697 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983 (1): 001–110 - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Roze,J.A. 1958. Los reptiles del Chimantá Tepui (Estado Bolívar, Venezuela) colectados por la expedición botánica del Chicago Natural History Museum. Acta Biol. Venezuelica 2 (25): 299-314
  • Señaris, J. Celsa; María Matilde Aristeguieta Padrón, Haidy Rojas Gil y Fernando J. M. Rojas-Runjaic 2018. Guía ilustrada de los anfibios y reptiles del valle de Caracas, Venezuela. Ediciones IVIC, Instituto Venezolano de Investigaciones Científicas (IVIC). Caracas, Venezuela. 348 pp.
  • Troschel, F.H. 1848. Amphibien. In: M.R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840-44. Im Auftrage Sr. Majestät des Königs von Preussen ausgeführt. Theil 3. Versuch einer Zusammenstellung der Fauna und Flora von British-Guiana. Leipzig, pp. 645-661 - get paper here
  • Vanzolini,P.E. & Williams,E.E. 1970. South American anoles: the geographic differentiation and evolution of the Anolis chrysolepis species group (Sauria, Iguanidae). Arquivos de Zoologia (São Paulo) 19: 1-298 - get paper here
  • Williams,E.E., Praderio,M.J. & Gorzula,S. 1996. A Phenacosaur from Chimanta Tepui, Venezuela. Breviora (506): 1-15 - get paper here
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