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Anolis smallwoodi SCHWARTZ, 1964

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
SubspeciesAnolis smallwoodi smallwoodi SCHWARTZ 1964
Anolis smallwoodi palardis SCHWARTZ 1964
Anolis smallwoodi saxuliceps SCHWARTZ 1964 
Common NamesE: Green-blotched Giant Anole, Smallwood's Anole 
SynonymAnolis equestris smallwoodi SCHWARTZ 1964: 412
Anolis smallwoodi — SCHWARTZ & GARRIDO 1972: 56
Anolis smallwoodi — SCHWARTZ & HENDERSON 1991: 343
Anolis smallwoodi — RODRÍGUEZ SCHETTINO 1999: 198
Anolis smallwoodi — NICHOLSON et al. 2005
Deiroptyx smallwoodi — NICHOLSON et al. 2012
Deiroptyx smallwoodi — NICHOLSON et al. 2018

Anolis smallwoodi smallwoodi SCHWARTZ 1964
Deiroptyx smallwoodi smallwoodi — NICHOLSON et al. 2012
Deiroptyx smallwoodi smallwoodi — NICHOLSON et al. 2018

Anolis smallwoodi palardis SCHWARTZ 1964
Anolis equestris palardis SCHWARTZ 1964: 416
Deiroptyx smallwoodi palardis — NICHOLSON et al. 2012
Deiroptyx smallwoodi palardis — NICHOLSON et al. 2018

Anolis smallwoodi saxuliceps SCHWARTZ 1964
Anolis equestris saxuliceps SCHWARTZ 1964: 422
Deiroptyx smallwoodi saxuliceps — NICHOLSON et al. 2012
Deiroptyx smallwoodi saxuliceps — NICHOLSON et al. 2018 
DistributionSE Cuba

smallwoodi: from Hongobosongo east to Bahia de Guantânamo; Sierra de la Gran Piedra. Type locality: Laguna de Baconao, Santiago de Cuba Province, Cuba.

palardis: southeastern Cuba from Guantânamo and Baitiquii inland to Sierra del Guaso.

saxuliceps: central southeastern Cuba; Type locality: Moa, Province Oriente , Cuba.  
TypesHolotype: AMNH 89526
Holotype: CM 33320, paratype: MCZ [palardis]
Holotype: ZMH (given as HM) 5376, paratypes: MCZ [saxuliceps] 
DiagnosisDESCRIPTION: Size large (SVL in males to 190 mm, in females to 165 mm);
moderate to high number (18-25) of vertical dorsals; size large to moderate (by population); metachrosis limited to shades of green in 1 population; dewlap pink, faintly orange, pale pink, or rosy; axillary stripe yellow to green; upper and lower jaws with prominent, pale labial stripe with green blotches on supralabials but without a subangular blotch; tail without broad, transverse bands. Dorsum green with paler green (at times orange) spotting, the spots encompassing as many as 6 adjacent scales, entire aspect leopard-like; occiput pale pea-green with bold, darker green vermiculations, or blotches and vermiculations absent (by population); casque black, gray, tan, brown, or blue, with or without pale dotting, flecking, or spotting; postlabial stripe pale green, bordered or not by darker green; eyelids at times bright orange or mustard; throat with darker (green?) longitudinally aligned dashes; venter pale green to green-brown; dewlap faint orange to pink; juveniles with 4 pale bands across body between limbs, first 2 often composed of dots, most posterior forming sacral U; transition to dotted dorsal pattern at SVL about 95-100 mm (Schwartz & Henderson 1991: 343). 
CommentSpecies group: Deiroptyx equestris species group (fide NICHOLSON et al. 2012). 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bona, Martin; Milan Novotný, Stanislav Danko and Adriana Burešová. 2012. Headstarting in a small population of European pond turtles (Emys orbicularis) in Central European conditions: first results. Herpetology Notes 6: 547-550
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos 2005. Mainland colonization by island lizards. Journal of Biogeography 32: 929–938 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. 1964. Anolis equestris in Oriente Province, Cuba. Bull. Mus. Comp. Zool. Harvard 131: 405-428. - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Garrido, O.H. 1972. The lizards of the Anolis equestris complex in Cuba. Stud. Fauna Curaçao and Carib. Is. 39 (134):1-86. - get paper here
  • Yuan, M. L., M. H. Wake, and I. J. Wang. 2019. Phenotypic integration between claw and toepad traits promotes microhabitat specialization in the Anolis adaptive radiation. Evolution 73: 231–244 - get paper here
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