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Aspidoscelis guttatus (WIEGMANN, 1834)

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Higher TaxaTeiidae, Teiinae, Gymnophthalmoidea, Sauria, Squamata (lizards)
SubspeciesAspidoscelis guttatus flavilineatus (DUELLMAN & WELLMAN 1960)
Aspidoscelis guttatus guttatus (WIEGMANN 1834)
Aspidoscelis guttatus immutabilis (COPE 1878) 
Common NamesE: Mexican Racerunner
S: Huico Ticuiliche Mexicano 
SynonymCnemidophorus guttatus WIEGMANN 1834: 29
Cnemidophorus unicolor COPE 1878: 93 (fide SMITH & TAYLOR 1950)
Cnemidophorus guttatus — GÜNTHER 1885: 28
Cnemidophorus guttatus guttatus — GADOW 1906: 320
Cnemidophorus guttatus — BURT 1931: 66 (part.)
Cnemidophorus guttatus guttatus — SMITH & TAYLOR 1950: 179
Cnemidophorus guttatus guttatus — SMITH & BURGER 1950
Cnemidophorus guttatus guttatus — MASLIN & SECOY 1986
Cnemidophorus guttatus — LINER 1994
Cnemidophorus guttatus — KÖHLER 2000: 100
Aspidoscelis guttata — REEDER et al. 2002
Aspidoscelis guttata guttata — LINER & CASAS-ANDREU 2008
Aspidoscelis guttata — MATA-SILVA et al. 2015
Aspidoscelis guttatus — PYRON & BURBRINK 2013

Aspidoscelis guttatus flavilineatus (DUELLMAN & WELLMAN 1960)
Cnemidophorus guttatus flavilineatus DUELLMAN & WELLMAN 1960
Cnemidophorus guttatus flavilineatus — KLUGE 1984
Cnemidophorus guttatus flavilineatus — MASLIN & SECOY 1986
Aspidoscelis guttata flavilineata — REEDER et al. 2002
Aspidoscelis guttata flavilineata — LINER & CASAS-ANDREU 2008

Aspidoscelis guttatus immutabilis (COPE 1878)
Cnemidophorus immutabilis COPE 1878: 93
Cnemidophorus guttatus immutabilis COPE 1892: 31
Cnemidophorus microlepidopus COPE 1877: 92
Cnemidophorus guttatus striatus GADOW 1903: 115
Cnemidophorus guttatus immutabilis — GADOW 1906: 320
Cnemidophorus immutabilis — WERNER 1910: 27
Cnemidophorus guttatus immutabilis — SMITH & TAYLOR 1950: 180
Cnemidophorus guttatus immutabilis — MASLIN & SECOY 1986
Aspidoscelis guttata immutabilis — REEDER et al. 2002
Cnemidophorus guttata immutabilis — CASTRO-FRANCO & BUSTOS-ZAGAL 2003
Aspidoscelis guttata immutabilis — LINER & CASAS-ANDREU 2008 
DistributionSE Mexico (Veracruz, Oaxaca, Guerrero through Chiapas)

Type locality: Mexico. Restricted to Veracruz, Veracruz, by SMITH & TAYLOR 1950.

flavilineatus: Mexico (Chiapas); Type locality: Mexico, Chiapas, Finca Orizaba, ca. 20 km SW Las Cruces, ca. 650 m elevation.

immutabilis: Mexico (Morelos)

striatus: Pacific slopes from Colima to Chiapas. Michoacán, Guerrero, Morelos. Type locality: “West Tehuantepec”, Oaxaca. Restricted to the city of Tehuantepec by SMITH & TAYLOR 1950.  
Reproductionoviparous 
TypesLectotype: ZMB 887 (designated by Burt, 1931:66, although given as 867, a lapsus calami; corrected by Duellman and Wellman, 1960:58); F. Deppe; 1824-1829
Holotype: USNM 30141; female; F. Sumichrast [immutabilis]
Holotype: UMMZ 119465; male; F. G. Thompson; 22 Dec. 1955 [flavilineatus]
Holotype: USNM 30385 [unicolor]
Holotype: USNM 30187 [Cnemidophorus microlepidopus] 
Diagnosis 
CommentSynonymy after SMITH & TAYLOR 1950. Belongs to the Aspidoscelis deppii group (fide REEDER et al. 2002).

Types: Malnate 1971 listed 2 syntypes (ANSP 9579-80) for Cnemidophorus guttatus Hallowell 1855: 192. 
EtymologyNamed after Latin guttatus, provided with drops, spots or specks.[“...Color superne cineraceus, guttis albidis (in vivo animali pallide viridibus?), in series quatuor longitudinales dispositis,

A. g. flavilineatus was named after Latin flavus, yellow, golden, gold colored + Latin lineatus, lined. ["...Frorn the Latin, flavus, yellow, and lineatus, lined; used in reference to the broad yellow middorsal stripe in adults…"].

A. g. immutabilis was named after Latin immutabilis, unchangeable/unalterable. ["...This species attains to the largest size known in the genus, without losing its striped coloration, as do the other large forms…"]. 
References
  • Alvarez DEL TORO, M. 1982. Los Reptiles de Chiapas. 3rd ed. México: Tuxtla Gutiérrez, 248 pp.
  • Bauer, A.M. and R. Günther 1994. An annotated type catalogue of the teiid and microteiid lizards in the Zoological Museum, Berlin (Reptilia: Squamata: Teiidae and Gymnophthalmidae). Mitt. Zool. Mus. Berlin 70:267-280 - get paper here
  • BELLO-SÁNCHEZ, EDGAR AHMED; JORGE E. MORALES-MÁVIL & ORLANDO R. VIVANCO-MONTANÉ. 2021. ASPIDOSCELIS GUTTATUS (Mexican Whiptail Lizard). PREDATION. Herpetological Review 52 (3): 644. - get paper here
  • Burt, C. E. 1931. A study of the teiid lizards of the genus Cnemidophorus with special reference to their phylogenetic relationships. Bull. US Natl. Mus. 154: 286 pp. - get paper here
  • Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F.
  • Castro Franco, R. & M. G. Bustos-Zagal 2003. Lagartijas de Morelos, México: distribución, hábitat y conservación. Acta Zoológica Mexicana (n.s.) 88:123-142 - get paper here
  • Castro-Franco, Rubén, María Guasalupe Bustos-Zagal 1994. List of reptiles of Morelos, Mexico, and their distribution in relation to vegetation types. Southwestern Naturalist 39 (2): 171-175 - get paper here
  • Cope, E.D. 1878. Tenth contribution to the herpetology of Tropical America. Proc. Amer. Philos. Soc. 17: 85-98 [1877] - get paper here
  • Cope, E.D. 1892. A synopsis of the species of the teiid genus Cnemidophorus. Trans. Am. Phil. Soc. (ser. 2) 17 (1): 27-52 [1893] - get paper here
  • Duellman, W. E. & WELLMAN, J. 1960. A systematic study of the lizards of the deppei group (genus Cnemidophorus) in México and Guatemala. Misc. Publ. Mus. Zool. Univ. Michigan (No. 111): 1-81 - get paper here
  • Gadow,H. 1903. Proc. Roy. Soc. London 72: 115
  • Gadow,H. 1906. A contribution to the study of evolution based upon the Mexican species of Cnemidophorus. Proc. Zool. Soc. London 1906: 277-375 (+ maps) - get paper here
  • Günther, A. C. L. G. 1885. Reptilia and Batrachia. Biologia Centrali-Américana. Taylor, & Francis, London, 326 pp. [published in parts from 1885-1902; reprint by the SSAR 1987] - get paper here
  • Johnson, Jerry D.; Vicente Mata-Silva, Elí García Padilla, and Larry David Wilson 2015. The Herpetofauna of Chiapas, Mexico: composition, distribution, and conservation. Mesoamerican Herpetology 2 (3): 272–329. - get paper here
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Lemos-Espinal JA, Smith GR 2020. A conservation checklist of the herpetofauna of Morelos, with comparisons with adjoining states. ZooKeys 941: 121-144 - get paper here
  • Lemos-Espinal JA, Smith GR, Pierce LJS, Painter CW 2020. The amphibians and reptiles of Colima, Mexico, with a summary of their conservation status. ZooKeys 927: 99-125 - get paper here
  • Malnate, Edmond V. 1971. A catalogue of primary types in the Herpetological Collections of the Academy of Natural Sciences, Philadelphia (ANSP). Proccedings of the Academy of Natural Sciences of Philadelphia 123 (9): 345-375 - get paper here
  • Maslin, T. & Secoy, D.M. 1986. A checklist of the lizard genus Cnemidophorus (Teiidae). Contr. Zool. Univ. Colorado Mus. 1: 1-60
  • Mata-González, Stephanye, Vicente Mata-Silva, Dominic L. DeSantis, Elí García-Padilla and Larry David Wilson. 2016. Aspidoscelis guttata (Wiegmann, 1834). Opportunistic water acquisition. Mesoamerican Herpetology 3 (2): 483-484 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
  • Palacios-Aguilar, Ricardo & OSCAR FLORES-VILLELA 2018. An updated checklist of the herpetofauna from Guerrero, Mexico. Zootaxa 4422 (1): 1-24 - get paper here
  • Peterson, A.T. et al. 2004. A preliminary biological survey of Cerro Piedra Larga, Oaxaca, Mexico: Birds, mammals, reptiles, amphibians, and plants. Anales del Instituto de Biología, Universidad Nacional Autónoma de México, Serie Zoología 75(2): 439-466
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • REEDER, T.W.; CHARLES J. COLE AND HERBERT C. DESSAUER 2002. Phylogenetic Relationships of Whiptail Lizards of the Genus Cnemidophorus (Squamata: Teiidae): A Test of Monophyly, Reevaluation of Karyotypic Evolution, and Review of Hybrid Origins. American Museum Novitates 3365: 1-64 - get paper here
  • Smith, H. M., & Burger, W.L. 1950. Herpetological results of the University of Illinois field expedition, Spring 1949. III. Sauria. Transactions of the Kansas Academy of Science 53 (2): 165-175 - get paper here
  • Torres-Hernández, LA, Ramírez-Bautista A, Cruz-Elizalde R, Hernández-Salinas U, Berriozabal-Islas C, DeSantis DL, Johnson JD, Rocha A, García-Padilla E, Mata-Silva V, Fucsko LA, and Wilson LD. 2021. The herpetofauna of Veracruz, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 15(2) [General Section]: 72–155 - get paper here
  • Walker, J M; Corneil, J P; Kartje, M E; Montgomery, C E & Shandersky, L; 2019. Aspidoscelis guttatus (Mexican Racerunner) Juvenile characteristics. Herpetological Review 50 (2): 364-365 - get paper here
  • Walker, J M; Kartje, M E; Corneil, J P & Montgomery, C E; 2019. Aspidoscelis guttatus (Mexican Racerunner) and Sceloporus siniferus (Long-tailed Spiny Lizard) Opportunistic foraging. Herpetological Review 50 (3): 568-569 - get paper here
  • Werner,F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Jahrb. Hamburg. Wiss. Anst., vol. 27 (1909), suppl. no. 2, 1910, pp. 1-46; reprinted: 1910, Mitteil. Naturhist. Mus. Hamburg, vol. 27: 205-) - get paper here
  • Wiegmann,A.F.A. 1834. Herpetologia Mexicana, seu descriptio amphibiorum novae hispaniae , quae itineribus comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede im Museum Zoologicum Berolinense Pervenerunt. Pars prima, saurorum species. Berlin, Lüderitz, iv + 54 pp. - get paper here
 
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