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Asthenodipsas ingeri QUAH, LIM & GRISMER, 2021

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Higher TaxaPareidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Inger’s Slug Snake 
SynonymAsthenodipsas ingeri QUAH, LIM & GRISMER 2021
Amblycephalus vertebralis – LOVERIDGE 1938: 43
Asthenodipsas vertebralis – STUEBING et al. 2014: 82 & 86 (part)
Pareas vertebralis – COX et al. 1998: 79 (part)
Pareas vertebralis – DE HAAS 1950: 529 (part)
Pareas vertebralis – HAILE 1958: 759, 766 (part)
Pareas vertebralis – ISKANDAR & COLIJN “2001” 2002: 113 (part)
Pareas vertebralis – MANTHEY & GROSSMANN 1997: 308, 378 (part)
Pareas vertebralis – STUEBING 1991: 331 (part)
Pareas vertebralis – STUEBING & INGER 1999: 87 (part)
Pareas vertebralis – INGER & VORIS 2001: 890 (part)
Amblycephalus laevis – SMITH 1925: 20 (part)
Asthenodipsas laevis – GROSSMANN & TILLACK 2003: 180 (part)
Asthenodipsas laevis – STUEBING et al. 2014: 82 (part)
Internatus laevis – MALKMUS et al. 2002: 342 (part)
Pareas laevis – STUEBING & INGER 1999: 87 (part)
Pareas carinatus – DAS 2012: 132 (part)
Pareas carinatus – DAS 2018: 132 (part)
Asthenodipsas (Asthenodipsas) ingeri — POYARKOV et al. 2022 
DistributionEast Malaysia (Borneo: Sabah and possibly Sarawak)

Type locality: Lumu Lumu, Mount Kinabalu, Sabah, East Malaysia (estimated: N 6.01966, E 116.539501, 1728 m a.s.l.).  
ReproductionOviparous (oresumably, Quah et al. 2021) 
TypesHolotype: MCZ R43592, Adult female, collected by John A. Griswold Jr. in July 1937 
DiagnosisDiagnosis: Asthenodipsas ingeri sp. nov. can be differentiated from its congeners by the following combination of characters: a maximum SVL of 715 mm; 15/15/15 dorsal scale rows; 201 ventrals; 53 subcaudal scales; one or two postoculars; 2+2+2 temporals; seven supralabials, 3rd,4th & 5th (sometimes 3rd & 4th) touching the eye; six to seven infralabials, 1st and 3rd pair in contact; single anterior inframaxillary present; two pairs of posterior inframaxillaries; a sharp vertebral keel; dorsum of adults brown to grey and overlain with up to 51 irregularly-shaped, rhomboidal dark-brown bands beginning on the nape and extending the length of body and tail and onto the lateral edges of the ventral scales to form spots, but not encircling body; a narrow, light-coloured vertebral stripe; throat and ventrals white to cream-coloured with very fine speckling; top of head and snout darker than ground colouration of body while the supralabials and temporal region are lighter in colour; and iris and pupils black (Table 2 & 3; fig. 2, 4 & 5 in Quah et al. 2021).


Additional details (2061 characters) are available for collaborators and contributors. Please contact us for details. 
CommentDiet: Presumably feeds on snails like other congeners (Quah et al. 2021)

Presumably lays eggs (Quah et al. 2021) 
References
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • Das, I. 2018. A Naturalist’s Guide to the Snakes of Thailand and South-east Asia. 2nd edition. John Beaufoy Publishing, Oxford, 176 pp
  • Grossmann, W. & Tillack, F. 2003. On the taxonomic status of Asthenodipsas tropidonotus (LIDTH DE JEUDE 1923) and Pareas vertebralis BOULENGER 1900) (Serpentes: Colubridae: Pareatinae). Russ. J. Herpetol. 10 (3): 175-190 - get paper here
  • HAAS, C.P.J. De 1950. Checklist of the snakes of the Indo-Australian archipelago. Treubia 20 (3): 511-625 - get paper here
  • Haile, N.S. 1958. The snakes of Borneo, with a key to the species. Sarawak Mus., Kuching, J. 8: 743-771.
  • Inger, R.F. & Voris, H. K. 2001. The biogeographical relations of the frogs and snakes of Sundaland. Journal of Biogeography 28: 863-891
  • Iskandar, D.T. & E. Colijn. 2001. Checklist of Southeast Asian and New Guinea Herpetofauna II Reptilia, Serpentes. Treubia 31:135–313 - get paper here
  • Loveridge, A. 1938. New snakes of the genera Calamaria, Bungarus and Trimeresurus from Mt. Kinabalu, North Borneo. Proc. Biol. Soc. Washington 51: 43-46. - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. (ISBN 3-931587-12-6) - get paper here
  • Poyarkov NA, Nguyen TV, Pawangkhanant P, Yushchenko PV, Brakels P, Nguyen LH, Nguyen HN, Suwannapoom C, Orlov N, Vogel G. 2022. An integrative taxonomic revision of slug-eating snakes (Squamata: Pareidae: Pareineae) reveals unprecedented diversity in Indochina. PeerJ 10: e12713 - get paper here
  • Quah, Evan S H; Kelvin K P Lim, L Lee Grismer 2021. On the taxonomic status of Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareidae) in Borneo with the description of a new species. Zootaxa 4949 (1): 24–44 - get paper here
  • Smith, M. A. 1925. Contribution to the herpetology of Borneo. Sarawak Mus. J. 3 (8): 15-34. - get paper here
  • Stuebing, R.B. 1991. A checklist of the snakes of Borneo. Raffles Bull. Zool. 39(2): 323-362. - get paper here
  • Stuebing, R.B. & INGER, R.F. 1999. A field guide to the snakes of Borneo. Natural history Publications (Borneo), Kota Kinabalu, 254 pp. [corrections in HR 31: 124]. - get paper here
  • Stuebing, R.B., Inger, R.F. & Lardner, B. 2014. A field guide to the snakes of Borneo, second edition. Natural history Publications (Borneo)
 
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