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Atractus savagei SALAZAR-VALENZUELA, TORRES-CARVAJAL & PASSOS, 2014

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Savage’s Groundsnake
S: Culebra Tierreras de Savage 
SynonymAtractus savagei SALAZAR-VALENZUELA, TORRES-CARVAJAL & PASSOS 2014 
DistributionEcuador (Carchi)

Type locality: surroundings of Chilmá Bajo on the way to Tres Marías waterfall, 0° 51’53.82’’N, 78°2’59.23’’W; datum 1⁄4 WGS84; 2071 m elevation), Tulcán County, Carchi Province, Ecuador.  
Reproduction 
TypesHolotype: QCAZ 8713 (Museo de Zoologí, Pontificia Universidad Cato ́lica del Ecuador), adult female, collected on 25 February 2009 by Omar Torres-Carvajal, Silvia Alda ́s-Alarco ́n, and Elicio E. Tapia; paratypes: QCAZ, MNRJ, 
DiagnosisDiagnosis: Atractus savagei is distinguished from all congeners by the following combination of characters: (1) 17–17–17 smooth dorsal scale rows; (2) two postoculars; () long loreal; (4) temporals 1 þ 2; (5) six supralabials, third and fourth contacting orbit; (6) seven infralabials, first four contacting chinshields; (7) 5–8 maxillary teeth; (8) four gular scale rows; (9) four preventrals; (10) 161–165 ventrals in females, 149–154 in males; (11) 2–26 subcaudals in females, 28– in males; (12) brown dorsum with black specks on the margins of the scales and two black longitudinal stripes on each side of the body: one occupying the upper and lower half of the first and second dorsal scale rows, respectively, and a second stripe present on the upper margin of the fourth dorsal scale row; (1) ventral ground color of head yellow, gradually changing to orange and red toward the back of the body, with two lateral and two medial series of round to rhomboidal black blotches arranged into four conspicuous longitudinal stripes; (14) moderate body size (females and males reaching 5 and 297 mm SVL, respectively); (15) long tail in males (14.4–18.2% SVL), moderate (11.–12.5% SVL) in females; (16) hemipenis moderately bilobed, barely capitate, and slightly calyculate.
 
CommentSize: max. 5 mm SVL, max. 50 mm caudal lenght, max. 85 total length 
EtymologyThe specific epithet is a noun in the genitive case, in honor of Professor Jay M. Savage for his lifetime of fruitful work with the New World herpetofauna (Donnelly, 2013), and also as a tribute to his seminal monograph on Ecuadorian Atractus (Donnelly, 2013). 
References
  • Donnelly, Maureen A. D 2013. Jay M. Savage. Copeia Dec 2013, Vol. 2013, No. 4: 757-767. - get paper here
  • Salazar-Valenzuela, David; Omar Torres-Carvajal, and Paulo Passos 2014. A New Species of Atractus (Serpentes: Dipsadidae) from the Andes of Ecuador. Herpetologica 70 (3): 350-363. - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
 
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