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Atractus werneri PERACCA, 1914

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Werner's Ground Snake 
SynonymAtractus werneri PERACCA 1914: 102
Atractus werneri — DUNN 1944: 21
Atractus werneri — PASSOS & LYNCH 2010
Atractus werneri — WALLACH et al. 2014: 84
Atractus werneri — PASSOS et al. 2024: 43 
DistributionColombia (Cundinamarca: Western slopes of Eastern Cordillera), elevation 1200-1800 m

Type locality: Viotá, Dep. de Cundinamarca, Colombia, 1830 m elevation, (04° 26’ N, 74° 31’ W).  
Reproductionoviparous 
TypesHolotype: MHNN, female (given erroneously as MNHN-RA 91.506 by some sources), given as NHNN by Passos et al. 
DiagnosisDiagnosis: Atractus werneri can be distinguished from all congeners by the following combination of characters: (1) dorsal scale rows smooth, 17/17/17; (2) postoculars 2; (3) loreal long; (4) temporal formula 1 + 2; (5) supralabials usually seven, 3 and 4 contacting eye; (6) infralabials usually seven, 1−3 contacting chinshields; (7) maxillary teeth 6−10; (8) gular scale rows three or four; (9) preventrals usually five; (10) ventrals 158−174 in females, 148−160 in males; (11) subcaudals 21−36 in females, 27−37 in males; (12) in preservative, dorsum uniformly brown or with conspicuous vertebral and dorsolateral lines; (13) in preservative, belly predominantly creamish white, occasionally with diffuse, irregular, dark brown blotches; (14) body size moderate, females reaching 346 mm SVL, males 330 mm SVL; (15) tail length short to moderate in females (8.7−15.2% SVL), moderate to long in males (13.2−17.1% SVL); (16) hemipenis slightly bilobed, semicapitate, semicalyculate; (17) laterosphenoid present, well developed. (Passos et al. 2024)


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CommentThe date of publication of Peracca’s paper is usually given as 1914. However, it has been shown that his paper was actually published at least in early 1913 but most likely in 1912.

Similar species: A. obtusirostris and A. melanogaster. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Dunn, Emmett R. 1944. Herpetology of the Bogota area. Revista de la Academia Colombiana de Ciencias Exacta, Fisicas y Naturales 6: 68-81
  • Passos, P., Meneses-Pelayo, E., Ramos, L. O., Martins, A. R., Machado, A., Lopes, R. T., ... & Lynch, J. D. 2024. Taxonomy without borders: Revision of the genus Atractus (Serpentes: Dipsadidae) from the Andes between Colombia and Venezuela. South American Journal of Herpetology, 32(sp1), 1-123 - get paper here
  • Passos, Paulo and John D. Lynch 2010. Revision of Atractus (Serpentes: Dipsadidae) from Middle and Upper Magdalena Drainage of Colombia. Herpetological Monographs 24 (1): 149-173 - get paper here
  • Peracca, M.G. 1914. Reptiles et Batraciens de Colombie. Pp. 96-111 In: Fuhrmann, O. and E. Mayor (eds.), Voyage d'exploration scientifique en Colombie. Memoires de la Societe neuchateloise des Sciences naturelles, 5
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Westersheim, Otto v. 1940. Franz Werner als Mensch und Forscher. Annalen des Naturhistorischen Museums in Wien 51: 8-53 - get paper here
 
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