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Atractus riveroi ROZE, 1961

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Rivero's Ground Snake 
SynonymAtractus riveroi ROZE 1961: 114
Atractus riveroi — PETERS & OREJAS-MIRANDA 1970: 34
Atractus riveroi — KORNACKER 1999: 65
Atractus riveroi — PASSOS et al. 2007
Atractus riveroi — WALLACH et al. 2014: 80
Atractus riveroi — NOGUEIRA et al. 2019 
DistributionVenezuela (TF Amazonas), Brazil (Roraima), elevation 1300-1800 m.

Type locality: Cerro Duida (1800 m elevation), Territorio Federal Amazonas, Venezuela, (03°25’N, 65°40’W.  
Reproductionoviparous 
TypesHolotype: AMNH 36615, adult male, collected on 20 April 1929 by members of the Tyler–Duida expedition; Paratype: MBUCV 7175, formerly UPR 49 (r-20) 
DiagnosisDiagnosis: Atractus riveroi can be distinguished from all congeners by the following combination of characters: (1) smooth dorsal scale rows 17/17/17; (2) postoculars two; (3) loreal long; (4) temporals 1 + 2; (5) supralabials eight, fourth and fifth contacting orbit; (6) infralabials eight, first four contacting chin shields; (7) maxillary teeth seven; (8) gular scale rows three or four; (9) preventrals three or four; (10) ventrals 152–161 in males; (11) subcaudals 34–41 in males; (12) in preservation, dorsum variable, ground color beige with dark brown transversal bands or ground color black with light transversal bands; (13) in preservation, venter cream with rhomboidal dark brown blotches linearly arranged forming irregular stripes; (14) body size moderate, males reaching 415 mm SVL; (15) tail relatively long in males (16.2–18.2% SVL); (16) hemipenis moderately bilobed, semicapitate, semicalyculate [PASSOS et al. 2013]. 
Comment 
EtymologyNamed after Dr. Juan Arturo Rivero-Quintero (b. 1923), who gained his bachelor's degree (1945) from Universidad de Puerto Rico, and both his master's (1951) and doctorate (1953) from Harvard. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • De Fraga, R., et al. 2017. Narrow endemism or insufficient sampling? Geographic range extension and morphological variation of the poorly known Atractus riveroi Roze, 1961 (Serpentes: Dipsadidae). Herpetological Review 48 (2): 281-284 - get paper here
  • Kornacker,P.M. 1999. Checklist and key to the snakes of Venezuela. PaKo-Verlag, Rheinbach, Germany, 270 pp.
  • Lancini,A.R. & Kornacker,P.M. 1989. Die Schlangen von Venezuela. Armitano Editores C.A., Caracas, 1-381
  • Moraes LJCL, Almeida AP, Fraga R, Rojas RR, Pirani RM, Silva AAA, de Carvalho VT, Gordo M, Werneck FP 2017. Integrative overview of the herpetofauna from Serra da Mocidade, a granitic mountain range in Northern Brazil . ZooKeys 715: 103-159 - get paper here
  • Natera-Mumaw, Marco; Luis Felipe Esqueda-González & Manuel Castelaín-Fernández 2015. Atlas Serpientes de Venezuela. Santiago de Chile, Dimacofi Negocios Avanzados S.A., 456 pp. - get paper here
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • Passos, P., R. Fernandes and Borges-Nojosa, D.M. 2007. A New Species of Atractus (Serpentes: Dipsadinae) from a Relictual Forest in Northeastern Brazil. Copeia 2007 (4): 788–797 - get paper here
  • Passos, Paulo; Philippe J. R. Kok, Nelson R. de Albuquerque, and Gilson A. Rivas 2013. Groundsnakes of the Lost World: A Review of Atractus (Serpentes: Dipsadidae) from the Pantepui Region, Northern South America. Herpetological Monographs 27 (1): 52-86 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Roze, Jánis A. 1961. El genero Atractus (Serpentes: Colubridae) en Venezuela. Acta Biologica Venezuelica 3 (7): 103-119
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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