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Bothriechis nubestris DOAN, MASON, CASTOE, SASA & PARKINSON, 2016

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Talamancan Palm-Pitviper 
SynonymBothriechis nubestris DOAN, MASON, CASTOE, SASA & PARKINSON 2016
Bothriechis nubestris — MASON et al. 2019 
DistributionCosta Rica (San José)

Type locality: San Isidro de El General, Province of San José, COSTA RICA; approximately 3000 m  
TypesHolotype: UTA R-9637 (Campbell and Lamar, 2004: fig. 97B), an adult female; collected in October 1973 by Peter Seigfried. Paratypes. All from province of San José, COSTA RICA: UTA R-2801, 2808, 2850, 7327, 7463, 9635, 9636, 10433, all from San Isidro de El General District; UCR 5727, 5728, 5758, 11151, 15420, 15423, 15424, 15432, all from San Gerardo de Dota; UCR 12356, from División, Páramo District. 
DiagnosisDiagnosis: (1) a medium-sized slender arboreal pitviper; (2) dorsum green with heavy black mottling; (3) iris blackish; (4) superciliary scales absent; (5) interrictals 22–29; (6) supraoculars thin, usually kidney-shaped; (7) intersupraoculars 6–10; (8) partial rows or two rows of irregular scales between suboculars and supralabials; (9) infralabials 9–12; (10) first dorsals usually 21 (75.9%); (11) second dorsals usually 21 (58.6%); (12) third dorsals usually 17 (85.7%); (13) ventrals 150–160; (14) subcaudals 52–64; (15) tail prehensile.
Specimens of Bothriechis nubestris differ from B. schlegelii and B. supraciliaris by lacking superciliary scales (present in B. schlegelii and B. supraciliaris). Bothriechis nubestris differs from all other Bothriechis species except B. nigroviridis by having green dorsal coloration with heavy black mottling and a blackish iris. Bothriechis nubestris differs from B. nigroviridis (see Table 1 for summary) by the combination of having 150–160 ventral scales (136–149 in B. nigroviridis), thin, often kidney-shaped supraoculars with a wide intersupraocular space (B. nigroviridis usually have wider supraoculars, never kidney-shaped, with narrow intersupraocular space), and higher average counts of interrictals, dorsals, and subcaudal scales than B. nigroviridis. 
CommentSimilar species: B. nigroviridis 
EtymologyThe specific epithet means ‘belonging to the clouds’. It is derived from the Latin noun nubes, –is, meaning cloud, and the Latin suffix –estris, meaning belonging to. This name alludes to the fact that this species inhabits cloud forests. The common name Talamancan Palm-Pitviper refers to its range in the Cordillera de Talamanca 
  • DOAN, TIFFANY M.; ANDREW J. MASON, TODD A. CASTOE, MAHMOOD SASA & CHRISTOPHER L. PARKINSON 2016. A cryptic palm-pitviper species (Squamata: Viperidae: Bothriechis) from the Costa Rican highlands, with notes on the variation within B. nigroviridis. Zootaxa 4138 (2): 271–290 - get paper here
  • Mason AJ; Grazziotin F; Zaher H; Lemmon A; Moriarty Lemmon E & Parkinson C 2019. Reticulate evolution in nuclear Middle America causes discordance in the phylogeny of palm-pitvipers (Viperidae: Bothriechis). Journal of Biogeography 46 (5): 833-844 - get paper here
  • OLIVEIRA-DALLAND, LUIS G.; LAURA R.V. ALENCAR, LEANDRO R. TAMBOSI, PAOLA A. CARRASCO, RHETT M. RAUTSAW, JESUS SIGALA-RODRIGUEZ, GUSTAVO SCROCCHI & MARCIO MARTINS. 2022. Conservation gaps for Neotropical vipers: Mismatches between protected areas, species richness and evolutionary distinctiveness. Biological Conservation 275(109750). - get paper here
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