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Bothrocophias myrringae ANGARITA-SIERRA, CUBIDES-CUBILLOS & HURTADO-GÓMEZ, 2022

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymBothrocophias myrringae ANGARITA-SIERRA, CUBIDES-CUBILLOS & HURTADO-GÓMEZ 2022
Bothrocophias microphthalmus — CAMPBELL & LAMAR 1989: 255
Bothrocophias microphthalmus — CAMPBELL & LAMAR 2004: Plate 474 
DistributionColombia (Cundinamarca)

Type locality: vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia, coordinates: N 4.660602778, W –73.88491667; elevation 2,761 m. a.s.l.  
Reproduction 
TypesHolotype. INSZ 0268, an adult male, collected by local people and brought to Francisco Javier Ruiz, staff of the INS, on 29th August 2020. (Figs 1, 6G–I, 7, 8A–C in Angarita-Sierra et al. 2022)
Paratypes. COLOMBIA [n=5; Fig. 8C–D]: Cundinamarca: Municipality of Guayabetal. Locality: unknown, IAvH-R 6877, coordinates N 4.17508, W –73.88117 (approximate to the town). Municipality of Choachí. Locality: Palo Alto, IAvH-R6840, coordinates N 4.61578, W –73.8904. Municipality of Fómeque. Locality: vereda de Coasavistá, INSV-SR-00365, coordinates N 4.495001, W –73.852056. — Meta: Municipality of El Calvario. Locality: unknown, INSVSR-0099, coordinates N 4.358925, W –73.71358889. Municipality of San Juanito. Locality: unknown, IAvH-R7045, coordinates N 4.457913889, W –73.67618889 (approximate to the town). 
DiagnosisDiagnosis: Bothrocophias myrringae sp. nov. can be distinguished from all its congeners by the following combination of characters: (1) 157–161 ventral scales in females, 152–155 ventral scales in males; (2) internasal scales in contact or separated by a single small scale (3) absence of canthorostral scales; (4) lacunolabial scale usually present; (5) hemipenial lobes slim and cylindrical, moderately capitate distally, weakly ornamented toward the apex with large and scarce calyces with weakly spinulate edges; (8) bifurcation point of the hemipenial lobes about 2–4 sudcaudal scales; (9) hemipenial body ornamented by numerous mesial spines that increase in size from the center to periphery of the hemipenial body and arranged in oblique lines; (10) in sulcate view, lateral and mesial spines of the hemipenial body variable in size; (11) sulcus spermaticus walls weakly defined; (12) usually more than 28 dark-brown bands and/or opposite or juxtaposed trapezoid-shaped blotches with paler centers dorsally; and (13) ventral surfaces of the tail with bright reddish or orange-reddish speckles with black spots without a regular pattern and heavily marked with dark pigment towards the base (Fig. 7). (Angarita-Sierra et al. 2022)


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Comment 
EtymologyNamed after the Latin translation of the Spanish nickname “Mirringa,” which means “pinch” or something very small. The word “Mirringa” was popularized by Rafael Pombo (1833–1912), a Colombian poet and writer who wrote a popular fable titled “Mirringa Mirronga. The name of the new species is in honor of the educator Myriam Sierra Guerrero (mother of the first author). 
References
  • Angarita-Sierra T, Cubides-Cubillos SD, Hurtado-Gómez JP 2022. Hidden in the highs: Two new species of the enigmatic genus of toadheaded pitvipers of the genus Bothrocophias. Vertebrate Zoology 72 971–996 - get paper here
  • Díaz-Flórez, Ronald Alessandro; Juan Timms-Rangel 2023. On the defensive behavior of the High-Andean Toad-headed Pitviper, Bothrocophias myrringae (Squamata: Viperidae), in central Colombia. Reptiles & Amphibians 30 (1): e18146 - get paper here
  • Kwet, A. 2023. Liste der im Jahr 2022 neu beschriebenen Reptilien. Elaphe 2023 (3): 48-73
 
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