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Brachyseps anosyensis (RAXWORTHY & NUSSBAUM, 1993)

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Higher TaxaScincidae, Scincinae, Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymAmphiglossus anosyensis RAXWORTHY & NUSSBAUM 1993
Amphiglossus anosyensis — GLAW & VENCES 1994: 315
Amphiglossus anosyensis — SCHMITZ et al. 2005
Amphiglossus anosyensis — MIRALLES et al. 2011 
DistributionMadagascar

Type locality: Ampamakiesiny Pass, 24°32’ S, 46°51’ E, 820 m elevation, 14 km northwest of Ranomafana-Sud, Tôlanaro (Fort Dauphin) Fivodronana, Toliara (Tulear) Province, Madagascar.  
Reproduction 
TypesHolotype: UMMZ 195746 (RAN 36337), adult male 
DiagnosisDiagnosis. A medium-sized Amphiglossus, 65 mm maximum SVL with 2830 longitudinal rows of scales around midbody (all the same size), 67-70 scales between mental scale and anal scale, 33-34 presacral vertebrae, forelimb when laid forwards reaches tympanum, 17-19 lamellae under fourth toe of the hindfoot. Coloration dorsally dark brown, with small circular yellow spots on head, body, and tail which are largest on anterior half of the body. Tail underside pale pink and unspotted. No other species of Amphiglossus has markings like A. anosyensis. The following species have similar scale counts and presacral vertebrae numbers, but can be separated from Amphiglossus anosyensis by the indicated characters:A. splendidus has a median pair of dorsal scales on the body which are enlarged laterallyand bold transverse stripes on the head and body (Brygoo,1985);A. frontoparietalis has bold transverse bands on the head and anterior portion of the body, the tail underside is spotted (Brygoo, 1980c), and the forelimbs extend past the tympanum, usually to the eye (Angel, 1942;Boulenger, 1889); A. melanurus is uniform brown above with each dorsal scale marked distally by a single dark spot, has 35-45 presacral vertebrae and 70-92 scales between mental and anal scale (Brygoo, 1984b); A. macrocercus is uniform brown above with two dark lateral spots on each scale, 35-39 presacral vertebrae and has 70-82 scales between mental and anal (Brygoo, 1984b); A. gastrotictus is uniform brown with fine dark spots and 35-36 presacral vertebrae (Brygoo, 1984b, RAXWORTHY & NUSSBAUM 1993: 333). 
Comment 
EtymologyNamed after the Anosy Mountains (after the Malagasy word “nosy” = island, “anosy” = in the island). 
References
  • Andreone, F. & Greer, A.E. 2002. Malagasy scincid lizards: descriptions of nine new species, with notes on the morphology, reproduction and taxonomy of some previously described species (Reptilia, Squamata: Scincidae). Journal of Zoology 258, 139-181 - get paper here
  • Crottini, Angelica; Janina Dordel, Jörn Köhler, Frank Glaw, Andreas Schmitz, Miguel Vences 2009. A multilocus phylogeny of Malagasy scincid lizards elucidates the relationships of the fossorial genera Androngo and Cryptoscincus. Molecular Phylogenetics and Evolution 53 (1): 345-350 - get paper here
  • Erens, Jesse; Aurélien Miralles, Frank Glaw, Lars Chatrou, Miguel Vences 2016. Extended molecular phylogenetics and revised systematics of Malagasy scincine lizards. Molecular Phylogenetics and Evolution - get paper here
  • Glaw ,F. & Vences, M. 1994. A Fieldguide to the Amphibians and Reptiles of Madagascar. Vences & Glaw Verlag, Köln (ISBN 3-929449-01-3)
  • MIRALLES, AURÉLIEN, ACHILLE P. RASELIMANANA, DOMOINA RAKOTOMALALA, MIGUEL VENCES & DAVID R. VIEITES 2011. A new large and colorful skink of the genus Amphiglossus from Madagascar revealed by morphology and multilocus molecular study. Zootaxa 2918: 47–67 - get paper here
  • Raxworthy C J. NUSSBAUM R A. 1993. Four new species of Amphiglossus from Madagascar (Squamata: Scincidae). Herpetologica 49 (3): 326-341. - get paper here
  • Schmitz, A.; M.C. Brandley; P. Mausfeld; M. Vences; F. Glaw; R.A. Nussbaum and T.W. Reeder 2005. Opening the black box: phylogenetics and morphological evolution of the Malagasy fossorial lizards of the subfamily “Scincinae”. Molecular Phylogenetics and Evolution 34: 118–133 (published online 2004) - get paper here
 
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