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Broadleysaurus major (DUMÉRIL, 1851)

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Higher TaxaGerrhosauridae (Gerrhosaurinae), Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Rough-scaled Plated Lizard
G: Braune Schildechse, Sudan-Schildechse 
SynonymGerrhosaurus major major DUMÉRIL 1851
Gerrhosaurus major DUMÉRIL 1851
Gerrhosaurus zanzibaricus PFEFFER 1889
Gerrhosaurus bottegoi DEL PRATO 1895
Gerrhosaurus major var. zechi TORNIER 1901
Gerrhosaurus bergi WERNER 1906
Gerrhosaurus grandis BOULENGER 1908
Gerrhosaurus major grandis — HEWITT 1910
Gerrhosaurus zechi SCHMIDT 1919: 519
Gerrhosaurus zechi — CAMP 1923: 342
Gerrhosaurus major zechi — LOVERIDGE 1936: 64
Gerrhosaurus major zechi — MERTENS 1938
Gerrhosaurus major bottegoi — PARKER 1942: 77
Gerrhosaurus major grandis — FITZSIMONS 1943: 280
Gerrhosaurus major grandis — LOVERIDGE 1955: 175
Gerrhosaurus major bottegoi — BROADLEY 1987
Gerrhosaurus major major — BROADLEY 1987
Gerrhosaurus major major — AUERBACH 1987: 118
Gerrhosaurus major major — BOYCOTT 1992
Gerrhosaurus major major — BROADLEY & HOWELL 1991: 18
Gerrhosaurus major — LARGEN & SPAWLS 2006: 43
Gerrhosaurus major zechi — CHIRIO & LEBRETON 2007
Gerrhosaurus major bottegoi — BARTS 2008
Gerrhosaurus major zechi — ULLENBRUCH et al. 2010
Gerrhosaurus major bottegoi — TRAPE, CHIRIO & TRAPE 2012
Broadleysaurus major — BATES et al. 2013
Broadleysaurus major bottegoi — NCBI TAXONOMY 2016
Broadleysaurus major — PIETERSEN et al. 2021 
DistributionE Kenya, Tanzania, Mozambique, Malawi, N Zambia,
Ethiopia, Eritrea, Togo, N Democratic Republic of the Congo (Zaire), Republic of South Africa, Swaziland, Botswana,
Zimbabwe, Republic of South Sudan (RSS), Central African Republic, Somalia, Uganda,
Cameroon, Nigeria, Benin, Ghana

bottegoi: Central African Republic, Somalia, Djibouti

zechi: Benin, Cameroon

Type locality: Zanzibar Island.
Type locality: Zululand [grandis]  
TypesSyntypes: MNHN-RA 2809 and MNHN-RA 6539
Holotype: ZMB 18814 [bergi]
Syntypes: ZMB 16608 [zechi] 
DiagnosisDiagnosis (genus): The monophyly of Broadleysaurus is established on the basis of a suite of nuclear and mitochondrial genetic characters (see above). This genus of terrestrial gerrhosaurids is differentiated from members of the genus Gerrhosaurus and the new genus described below for ‘G. validus’ by the following morphological characters: Body stout, cyclotetragonal to feebly depressed dorso-ventrally, and well armoured; readily distinguished from the above-named genera by its much lower (31–38 versus 49–67) numbers of transverse dorsal scale rows; also distinguished by having 9–10 longitudinal rows of ventrals (8 in Gerrhosaurus, but 10 in G. typicus; 12–20 in the new genus described in Bates et al. 2013 for ‘G. validus’) (Loveridge 1942; FitzSimons 1943, 1953; De Witte 1953; Laurent 1954, 1964; Broadley 1966; De Waal 1978; Jacobsen 1989) [BATES et al. 2013]. 
CommentSynonymy: BATES et al. (2013) synonymized Gerrhosaurus (major) bottegoi with G. (now Broadleysaurus) major.

NCBI taxonomy: 219621 (bottegoi)

Type species: Gerrhosaurus major DUMÉRIL 1851 is the type species of the genus Broadleysaurus BATES & TOLLEY 2013. 
EtymologyApparently named after its large size or its major position within the genus.

The genus has been named in honour of Dr Donald George Broadley (1932-2016), Curator Emeritus at the Natural History Museum, Bulawayo (Zimbabwe) for his numerous contributions to African herpetology (e.g. see Branch 1997). He was ranked as the third most productive living author of reptile names (Uetz 2010). The name Broadleysaurus is masculine, derived from Broadley + the Latinised word saurus meaning lizard. See Bates & Branch 2018 for an obituary. 
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