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Bronchocela hayeki (MÜLLER, 1928)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Sumatra Bloodsucker 
SynonymCalotes hayeki MÜLLER 1928
Calotes hayeki — BRONGERSMA 1931
Calotes hayeki — WERMUTH 1967: 38
Bronchocela hayeki — MOODY 1980
Bronchocela hayeki — MANTHEY & GROSSMANN 1997: 161
Bronchocela hayeki — TEYNIÉ et al. 2010
Bronchocella hayeki — NUGRAHA et al. 2020 (in error) 
DistributionIndonesia (N Sumatra)

Type locality: Hochfläche von Brastag, am Sibayak, Nordost-Sumatra (elevation 1400 m). Neotype locality: near Berastagi 3°11’ N, 98° 31’ E, “Fuss des Sibayak” about 1450 m a.s.l., Sumatera Utara.  
Reproductionoviparous 
TypesNeotype. ZMB 55931 near Berastagi 3°11’ N, 98° 31’ E, “Fuss des Sibayak” about 1450 m a.s.l., Sumatera Utara, leg. Ulrich Manthey, April 1996, adult male (designated by Hallermann 2005). Holotype formerly ZSM 3/1928, destroyed fide Glaw, cited in Hallermann 2005. Other specimens: RMNH, ZMH, MZB, UIMZ 
DiagnosisDiagnosis. Bronchocela hayeki can be distinguished from its sympatric congener B. cristatella by the presence of larger gular scales and larger ventral scales in 8 rows (vs. 10 – 12 in cristatella) and crescent-shaped erect nuchals as long as or longer than diameter of orbit in males and a dark brown tympanum and dark colored orbit; from jubata and orlovi with similar nuchal scales and shape of gular sac by higher number of scales around midbody (58 – 75 vs. 43 – 55), from celebensis by a larger diameter of tympanum (more than half diameterof orbit). It differs from B. marmorata by more dorsal scale rows directed upward (7 – 9 vs. 1 – 2 in marmorata, jubata, orlovi, and 3 – 4 in celebensis). It differs from B. danieli, smaragdina and Bronchocela sp. nov. by fourth finger not longer than fifth toe.


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CommentDistribution: See map in Amarasinghe et al. 2022: 411 (Fig. 1) 
EtymologyNamed after Hans von Hayek (1869-1940), an Austrian artist who moved to Munich (1891) and traveled widely within Europe and Asia. 
References
  • Amarasinghe AAT, Kamsi M, Riyanto A, Putra CA, Hallermann J, Andayani N, Abinawanto A, Supriatna J. 2022. Taxonomy, distribution, and conservation status of a rare arboreal lizard, Bronchocela hayeki (Mller, 1928) (Reptilia: Agamidae) from northern Sumatra, Indonesia. Zootaxa 5120 (3):409-422 - get paper here
  • AMARASINGHE, ; A.A. THASUN; IVAN INEICH, AWAL RIYANTO, JAKOB HALLERMANN, NOVIAR ANDAYANI, A. ABINAWANTO, JATNA SUPRIATNA 2022. Taxonomy and distribution of a common arboreal lizard, Bronchocela jubata Duméril & Bibron, 1837 (Reptilia: Agamidae), with designation of its lectotype from Java, Indonesia. Zootaxa 5150 (1): 065–082 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • GRISMER, L. LEE; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK W. SITES, JR. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023 - get paper here
  • Hallermann, J. 2005. Mit Hörnern, Kämmen und Gleithäuten - die bizarren Baumagamen. Reptilia (Münster) 10 (51): 18-25 - get paper here
  • Hallermann, J. 2005. A taxonomic review of the genus Bronchocela (Squamata: Agamidae) with description of a new species from Vietnam. Russ. J. Herpetol. 12 (3): 167-182 - get paper here
  • Hallermann, J. 2005. The bizarre arboreal agamids. Reptilia (GB) (42): 8-15 - get paper here
  • Malkmus,R. & Kunkel, M. 1992. Amphibien und Reptilien aus dem Leuser-Nationalpark/Nordsumatra. Herpetofauna 14 (76): 27 - get paper here
  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Moody,S.M. 1980. Phylogenetic and historical biogeographical relationships of the genera in the family Agamidae (Reptilia: Lacertilia). PhD thesis, Univ. Michigan, Ann Arbor, 373 pp.
  • Müller, L. 1928. Herpetologische Mitteilungen II. Ein neuer Calotes von Sumatra. Zool. Anz. 77: 67-69
  • Nugraha, F. A. D., Selaras, G. H., & Satria, R. 2020. Preliminary Checklist of Herpetofauna of Mount Sago Along the Hiking Trail in the Dry Season. International Conference on Biology, Sciences and Education (ICoBioSE 2019) (pp. 51-55). Atlantis Press - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
 
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