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Bronchocela cristatella (KUHL, 1820)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Green crested lizard
G: Borneo-Langschwanzagame, Borneo-Schönechse, Blutsaugeragame 
SynonymAgama cristatella KUHL 1820: 108
Agama gutturosa MERREM 1820
Bronchocela cristatella — KAUP 1827: 619
Bronchocela gutturosa — KAUP 1827: 619
Calotes gutturosa — CUVIER 1829: plate 7
Agama moluccana LESSON 1830: 34
Bronchocela cristatella — DUMÉRIL & BIBRON 1837: 395
Pseudocalotes archiducissae FITZINGER 1861 (nom. nud. fide SMITH 1935)
Calotes (Bronchocele) philippinus PETERS 1867: 16 (part.)
Bronchocela intermedia PETERS & DORIA 1878: 375
Calotes cristatellus — BOULENGER 1885: 316
Calotes cristatellus — FLOWER 1899
Calotes octospinosus BAUMANN 1913
Calotes cristatellus — DE ROOIJ 1915: 121
Calotes cristatellus — TAYLOR 1922: 139
Calotes cristatellus — SMITH 1935: 184
Calotes cristatellus moluccanus — BRONGERSMA 1948: 489
Calotes cristatellus — TAYLOR 1963: 886
Calotes cristatellus — HENDRICKSON 1966: 64
Bronchocela cristatella — MANTHEY 1983: 25
Bronchocela petersdoriai BÖHME & BISCHOFF 1984 (nom. nov. pro B. intermedia)
Calotes cristatellus — BONG HEANG 1987
Calotes cristatellus — BROWN et al. 1996
Bronchocela cristatella — MANTHEY & GROSSMANN 1997: 160
Bronchocela cristatella — COX et al. 1998: 98
Bronchocela cristatella — GAULKE 1999
Bronchocela christatella — CHAN-ARD et al. 1999 (in error)
Bronchocela cf. cristatella — GAULKE 2001
Bronchocela cristatella — BARTS & WILMS 2003
Bronchocela cristatella — NUR-AMALINA et al. 2017
Bronchoecella cristatella — BADLI-SHAM et al. 2019 (in error)
Bronchocela cf. cristatella — GLÄSSER-TROBISCH & TROBISCH 2021 
DistributionMalaysia (Pulau Pangkor, Pulau Tioman, Pulau Langkawi, Pulau Singa Besar, Johor: Pulau Sibu, Borneo),
S Myanmar (= Burma) (Tenasserim), S Thailand, W Malaysia, Singapore, India (Andaman and Nicobar Islands), Philippines (Palawan, Calamian Islands, Panay, Luzon, Mindanao, Sulu, Cebu, Bohol, Leyte, Negros, Samar, Mindoro, Masbate), Indonesia (Halmahera, Sumatra, Morotai etc.), Papua New Guinea

Type locality: not given by KUHL.

Type locality: “Kandari nel S.E. di Selebes” [= Kendari, Sulawesi, Indonesia] [Bronchocela intermedia PETERS & DORIA 1878].  
TypesHolotype: lost fide Brygoo 1988 but Das & Gemel 2000 suggested that KUHL based his name on MNHN 6884 (three syntypes).
Holotype: NMW 21002: 1-2 [archiducissae] 
DiagnosisDiagnosis (genus): Bronchocela species are slender, medium sized (SVL about 80–140 mm) with long tails TL/SVL 3,2–3,9, laterally compressed body and relatively long legs. The genus can be separated from all other draconinae genera by a fold of skin that extends from the angle of the jaw to the insertion of the upper arm; homogeny lateral scales rhombic and small, point backwards and downwards, ventral scales greater; tympanum naked; nuchal crest present, lower dorsal crest sometimes reduced to a serrated ridge; spines on the head absent (U. Manthey, pers. comm., 3 Aug 2021).

Diagnosis. A variable species of 70.8 - 114.0 mm SVL (in examined material, mean = 88.77, n = 64, maximal SVL 120 mm; Diong and Lim, 1998) and a long tail ratio = 295 - 394 (336.02, n = 50). 8 - 10 supralabials and 7 - 12 infralabials. Tympanum large, more than half diameter of orbit, often dusky colored. Canthus rostralis sharp, a series of 3 - 5 small erect, compressed scales behind supraciliary edge forming a small longitudinal crest. No enlarged scales between tympanum and eye. Gular sac small in males and sometimes absent in females, covered with small keeled scales, smaller than ventrals. Nuchal crest scales small, erect, flattened and directed upwards, longest shorter than diameter of orbit. Dorsal crest formed as a serrated ridge continues nuchal crest or absent. Body scales mucronate, keeled, homogenous in 53 - 95 midbody scales (mean = 71.58, n = 55), 4 - 10 uppermost scale row next to dorsal crest pointing upward, 0 - 4 rows directed parallel, others scale rows downwards. Ventrals 1 - 5 times larger than dorsals, strongly keeled. It can be distinguished from other congeners by combination of short nuchal crest, 4 - 10 upper scale rows pointing upwards and 10 - 12 ventral scale rows. 
CommentDistribution: According to SMITH 1935 not on New Guinea.

Type species: Agama cristatella KUHL 1820 is the type species of the genus Bronchocela KAUP 1827: 619.

Synonymy: Boulenger (1880) synonymized B. burmana with Calotes cristatellus but Zug et al. 2017 revalidated Bronchocela burmana again. Bronchocela intermedia PETERS & DORIA 1878 is a junior homonym of B. intermedia BERTHOLD 1842.

The Bronchocela cristatella mentoned in ter Borg (2005) is probably B. jubata (ter Borg 2007). 
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