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Calorodius thorntonensis (GREER, 1983)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Thornton Peak Calyptotis, Thornton Peak Skink 
SynonymCalyptotis thorntonensis GREER 1983: 47
Calyptotis thorntonensis — COGGER 2000: 389
Calyptotis thorntonensis — WILSON & SWAN 2010
Calorodius thorntonensis — TORKKOLA et al. 2022 
DistributionAustralia (Queensland)

Type locality: southern base of Thornton Peak, Qld., 16° 11' S, 145°24' E.  
Reproductionoviparous 
TypesHolotype: QM J28354 (previously AMS R56574) 
DiagnosisDiagnosis (genus). A genus of Australian Sphenomorphini displaying the following combination of derived character states: head tapers to a point in profile, the frontals and nasals lying in the same plane which is angled down from the frontoparietal suture to the premaxilla; limbs short, hind limb length no more than 27% of SVL, toes short, with loss of two phalanges from the fourth finger of the manus (2.3.4.3.3) and one from the fourth toe of the pes (2.3.4.4.4); postorbital bone absent; small posteriorly-directed processes on the palatal rami of the pterygoids which are separated by deeply projecting posterior processes of the palatines; single loreal scale; lower secondary temporal scale is overlapped by the upper secondary temporal; postmental scale contacts only the first infralabial scale; midbody scale rows 22–24; external ear represented by scaly, discoidal depression (scaly, superficial tympanum). (Torkkola et al. 2022, supplementary file S7).

Diagnosis (species). Calyptotis thorntonensis differs from its congeners in each of the following characters: a more depressed head and body; external ear represented by a scaly, discoidal depression (scaly, superficial tympanum) instead of either a scaleless tympanum or a scaly conical depression, and generally more longitudinal scale rows at midbody (24 vs modes of 22 or 20). (Greer 1983)

Comparison with other Calyptotis. Comparisons between C. thorntonensis and C. ruficauda, C. lepidorostrum, C. scutirostrum and C. temporalis have been made on pp. 34, 39,44, and 47, respectively and also in Table 5 (GREER 1983: 48). 
CommentAbundance: only known from the type locality (Meiri et al. 2017).

Limb morphology: 5 digits, 5 toes (Singhal et al. 2018, Cogger 2014)

Type species: Calyptotis thorntonensis GREER 1983: 47 is the type species of the genus Calorodius Torkkola et al. 2022. 
EtymologyNamed after Thornton Peak, the type and, as yet, only known locality for the species.

The genus was named after the Latin masculine word for heat, calor, and for disliking, odio, in reference to its occurrence in relatively cool, high elevation habitats, and apparent physiological intolerance for heat. 
References
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Couper, P., Covacevich, J., Amey, A. & Baker, A. 2006. The genera of skinks (Family Scincidae) of Australia and its island territories: diversity, distribution and identification. in: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (eds.). Evolution and Zoogeography of Australasian Vertebrates. Australian Scientific Publishing, Sydney, pp. 367-384
  • Greer A E 1983. The Australian scincid lizard genus Calyptotis de Vis: resurrection of the name, description of four new species, and discussion of relationships. Rec. Austral. Mus. 35 (1): 29-59 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Shea, Glenn M; Sadlier, Ross A 1999. A catalogue of the non-fossil amphibian and reptile type specimens in the collection of the Australian Museum: types currently, previously and purportedly present. TECHNICAL REPORTS OF THE AUSTRALIAN MUSEUM 15, 1999: 1-91 - get paper here
  • Singhal, Sonal; Huateng Huang, Maggie R. Grundler, María R. Marchán-Rivadeneira, Iris Holmes, Pascal O. Title, Stephen C. Donnellan, and Daniel L. Rabosky 2018. Does Population Structure Predict the Rate of Speciation? A Comparative Test across Australia’s Most Diverse Vertebrate Radiation. The American Naturalist - get paper here
  • Skinner, Adam; Mark N. Hutchinson, Michael S.Y. Lee 2013. Phylogeny and Divergence Times of Australian Sphenomorphus Group Skinks (Scincidae, Squamata). Molecular Phylogenetics and Evolution 69 (3): 906–918 - get paper here
  • Torkkola, J. J., Wilmer, J. W., Hutchinson, M. N., Couper, P. J., & Oliver, P. M. 2022. Die on this hill? A new monotypic, microendemic and montane vertebrate genus from the Australian Wet Tropics. Zoologica Scripta 51, 483– 497 - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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