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Calotes bachae HARTMANN, GEISSLER, POYARKOV, IHLOW, GALOYAN, RÖDDER & BÖHME, 2013

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymCalotes bachae HARTMANN, GEISSLER, POYARKOV, IHLOW, GALOYAN, RÖDDER & BÖHME 2013
Calotes bachae — WAGNER et al. 2021 
DistributionS Vietnam (Dong Nai, Binh Phuoc), Cambodia

Type locality: Vietnam, Dong Nai Province, Cat Tien National Park (11°38'04'' N 107°27'24'' E), 104 m elevation.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZFMK 88935, an adult male (Fig. 4A), collected 10 May 2009 by PG. Paratypes: An adult female (IEBR A.2012.23; Fig. 4B) and a juvenile (ZFMK 88937; Fig. 4E), same data as holotype (except collection dates: 20 April 2009 (IEBR A.2012.23) and 14 June 2009 (ZFMK 88937)) by PG. Two adult males (ZMMU NAP-02910-911; Fig. 4D) from Phuoc Hoa environs, Bu Gia Map National Park, Dak Ka River valley, Bu Gia Map commune, Bu Gia Map District, Binh Phuoc Province, southern Vietnam (12°19.6’ N, 107°25.2’ E; ca. 400 m above sea level), collected on 26 May 2011 by NAP. Two adult females (ZMMU NAP- 01509, 01512) from Ma Da forestry, Dong Nai Nature Reserve (former Vinh Cuu Nature Reserve), Dong Nai Province, southern Vietnam, 80 m elevation, collected 12.06.2011 by NAP. 
DiagnosisDiagnosis: A medium-sized Calotes with a maximum SVL of 97 mm, distinguished from all congeners by the combination of the following characters: 1) head and body robust; 2) body scales relatively small, homogeneous, feebly keeled, arranged in regular rows; 3) upper dorsolateral scales pointing backwards and upwards; 4) 44–50 midbody scale rows; 5) two short and well separated spines, surrounded by three to four scales on either side of the upper head above tympanum; 6) nuchal and dorsal crest continuous, composed of erect compressed scales, directed posteriorly; 7) vertebral spines and scales 35–42 in males and 43–46 in females; 8) oblique fold of skin in front of forelimb insertion dinstict, covered with small granular dark scales; 9) extremities and tail relatively long and slender (see comparison with C. mystaceus for details); 10) bluish to turquoise head and anterior body part, this coloration not well exceeding front limb insertion; 11) yellowish light stripe at upper lip reaching from below anterior corner of eye to posterior end of head; 12) very faint or no dorsal brownish blotches, medially extending across the vertebral crest.


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CommentIllustration: Iguana 27 (2): 10 (2014).

Behavior: diurnal, semiarboreal 
EtymologyNamed after Rike Bach (Bonn), in gratitude for supporting the second author (PG) during several fieldtrips in Indochina. 
References
  • Geißler, Peter G 2013. Eine neue Schönechse aus Indochina. Terraria-Elaphe 2013 (5): 46 - get paper here
  • Hartmann, Timo, Peter Geissler, Nikolay A. J. Poyarkov, Flora Ihlow, Eduard A. Galoyan, Dennis Rödder & Wolfgang Böhme. 2013. A new species of the genus Calotes Cuvier, 1817 (Squamata: Agamidae) from southern Vietnam. Zootaxa 3599 (3): 246–260 - get paper here
  • Karthik, Pandi; Ayuthavel Kalaimani, Rathinalingam Nagarajan 2018. An inventory on herpetofauna with emphasis on conservation from Gingee Hills, Eastern-Ghats, Southern India. Asian Journal of Conservation Biology 7 (1): 2-16 - get paper here
  • Neang T., Grismer, L.L., Hun S. & Phan C. 2015. New herpetofauna records and range extensions for Daboia siamensis (Smith, 1917) and Gekko petricolus Taylor, 1962 from Cambodia. Cambodian Journal of Natural History, 2015: 172–182 - get paper here
  • Wagner, P., Ihlow, F., Hartmann, T., Flecks, M., Schmitz, A., & Böhme, W. 2021. Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141–171 - get paper here
  • Ziegler, T & Nguyen, QT 2015. Neues von den Forschungs- und Naturschutzprojekten in Vietnam und Laos. Zeitschrift des Kölner Zoos 2015 (2): 79-
  • Ziegler, Thomas & Anna Rauhaus 2015. Erste herpetologische Amtshilfe aus Köln im Zoo Saigon. Terraria-Elaphe 2015 (4): 72-80 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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