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Calotes irawadi ZUG, BROWN, SCHULTE & VINDUM, 2006

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymCalotes irawadi ZUG, BROWN, SCHULTE & VINDUM 2006
Calotes irawadi — GOWANDE et al. 2021
Calotes irawadi — LIU et al. 2021 
DistributionMyanmar (Burma), China (W Yunnan), India (Manipur)

Type locality: MYANMAR: Sagaing Division, Chatthin Wildlife
Sanctuary, San Myaung Camp (23°34′27.6′′N, 95°44′15.6′′E; ca. 110 m elevation), approx. 2 km WNW of Chatthin.  
Reproductionoviparous 
TypesHolotype: USNM 520543, an adult male by local collector, 17 July 1997. 
DiagnosisDiagnosis: Calotes irawadi is a member of the versicolor species group and shares the obliquely posteriorad or vertical scale-row orientation on the neck and adjacent supra-axillary area with all other versicolor group members except C. htunwini. It differs from C. versicolor (Pondicherry population) by a much smaller body size (female, male means 77.4, 82.4 mm SVLvs. 93 mm, 119 mm, respectively), and more Dorsal (means 48.9 vs. 40.8) and Midbody (47.1 vs 42.8). Middorsal crest scales in C. irawadi are smaller (equal to tympanum diameter) and are straighter and more numerous than C. versicolor (Pondicherry) with crest scales to 2.5–3.0×diameter of tympanum. Lengths of supratympanic spines in C. irawadi are half or less the diameter of tympanum and 2/3 or more tympanum diameter in C. versicolor (Pondicherry). C. irawadiaverages smaller than Moyingyi and Nat-Ma-Taung “versicolor” and has fewer Dorsal. It differs from eastern Thailand “versicolor” by more Dorsal and in coloration by the usual presence of DorsalSt and NucSpot [from ZUG & VINDUM 2006]. 
CommentDistribution: see map with localities in Gowande et al. 2021: 672 (Fig. 1) 
EtymologyIrawadi is a variant spelling of Ayeyarwaddy and is used as a noun in apposition. Our use of Irawadi refers to the broad distribution of this species in the central portion of the Ayeyarwaddy River basin. 
References
  • Das, Abhijit; Uttam Saikia, B. H. C. K. Murthy, Santanu Dey and Sushil K. Dutta 2009. A herpetofaunal inventory of Barail Wildlife Sanctuary and adjacent regions, Assam, north-eastern India. Hamadryad 34 (1): 117 – 134 - get paper here
  • Decemson, H., Sinate, R. R., Biakzuala, L., Mathipi, V., Lalnuntluanga, P., Malsawma, J., & Lalremsanga, H. T. 2023. Confirmation of Irawadi Forest Lizard, Calotes irawadi Zug, Brown, Schulte and Vindum 2006 (Squamata: Agamidae), from Manipur, Northeast India. Reptiles & Amphibians, 30(1): e17043-e17043 - get paper here
  • Gowande G, Pal S, Jablonski D, Masroor R, Phansalkar PU, Dsouza P, Jayarajan A, Shanker K 2021. Molecular phylogenetics and taxonomic reassessment of the widespread agamid lizard Calotes versicolor (Daudin, 1802) (Squamata, Agamidae) across South Asia. Vertebrate Zoology 71: 669-696 - get paper here
  • Huang, Y., Li, H., Wang, Y., Li, M., Hou, M., & Cai, B. 2023.  Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys, 1187, 63-89 - get paper here
  • Liu S, Zuo C, Rao D 2021. Distribution extension of Calotes irawadi Zug, Brown, Schulte & Vindum, 2006, previously confused with C. versicolor (Daudin, 1802): first record from China. Herpetozoa 34: 83-88 - get paper here
  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
  • SÁNCHEZ-VIALAS, A.; COPETE-MOSQUERA, L. A., & CALVO-REVUELTA, M. 2024. Contributions to the amphibians and reptiles of Myanmar: insights from the Leonardo Fea legacy housed at the Museo Nacional de Ciencias Naturales of Madrid. Zootaxa, 5457(1), 1-64 - get paper here
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Zug, G. R., H. H. K. Brown, J. A. Schulte II, and J. V. Vindum 2006. Systematics of the Garden Lizards, Calotes versicolor Group (Reptilia, Squamata, Agamidae), in Myanmar: Central Dry Zone Populations. Proc. Cal. Acad. Sci. 57 (2):35-68. - get paper here
 
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