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Calotes bachae HARTMANN, GEISSLER, POYARKOV, IHLOW, GALOYAN, RÖDDER & BÖHME, 2013

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymCalotes bachae HARTMANN, GEISSLER, POYARKOV, IHLOW, GALOYAN, RÖDDER & BÖHME 2013
Calotes bachae — WAGNER et al. 2021 
DistributionS Vietnam (Dong Nai, Binh Phuoc), Cambodia

Type locality: Vietnam, Dong Nai Province, Cat Tien National Park (11°38'04'' N 107°27'24'' E), 104 m elevation.  
Reproduction 
TypesHolotype: ZFMK 88935, an adult male (Fig. 4A), collected 10 May 2009 by PG. Paratypes: An adult female (IEBR A.2012.23; Fig. 4B) and a juvenile (ZFMK 88937; Fig. 4E), same data as holotype (except collection dates: 20 April 2009 (IEBR A.2012.23) and 14 June 2009 (ZFMK 88937)) by PG. Two adult males (ZMMU NAP-02910-911; Fig. 4D) from Phuoc Hoa environs, Bu Gia Map National Park, Dak Ka River valley, Bu Gia Map commune, Bu Gia Map District, Binh Phuoc Province, southern Vietnam (12°19.6’ N, 107°25.2’ E; ca. 400 m above sea level), collected on 26 May 2011 by NAP. Two adult females (ZMMU NAP- 01509, 01512) from Ma Da forestry, Dong Nai Nature Reserve (former Vinh Cuu Nature Reserve), Dong Nai Province, southern Vietnam, 80 m elevation, collected 12.06.2011 by NAP. 
DiagnosisDiagnosis: A medium-sized Calotes with a maximum SVL of 97 mm, distinguished from all congeners by the combination of the following characters: 1) head and body robust; 2) body scales relatively small, homogeneous, feebly keeled, arranged in regular rows; 3) upper dorsolateral scales pointing backwards and upwards; 4) 44–50 midbody scale rows; 5) two short and well separated spines, surrounded by three to four scales on either side of the upper head above tympanum; 6) nuchal and dorsal crest continuous, composed of erect compressed scales, directed posteriorly; 7) vertebral spines and scales 35–42 in males and 43–46 in females; 8) oblique fold of skin in front of forelimb insertion dinstict, covered with small granular dark scales; 9) extremities and tail relatively long and slender (see comparison with C. mystaceus for details); 10) bluish to turquoise head and anterior body part, this coloration not well exceeding front limb insertion; 11) yellowish light stripe at upper lip reaching from below anterior corner of eye to posterior end of head; 12) very faint or no dorsal brownish blotches, medially extending across the vertebral crest.

See also Wagner et al. 2021 for comparisons.

Coloration (males). The brilliant coloration of adult males is characterized by a brightly colored bluish to tur- quoise head, with bluish coloration continuing posterior- ly to fore limb insertion. There is a relatively faded light (bright yellowish, when under acute distress, a character typical for C. bachae only) stripe at the upper lip crossing the tympanum from beneath the eye to end of head. Gular pouch colored in darker blue, interscale skin black. A tri- angular to crescent-shaped patch of small black scales is present in front of shoulder. Very faint brownish blotch- es extending mid-dorsally over the vertebral crest, from above fore limb insertion on to tail; posterior to fore limb insertion brownish orange in color on trunk and tail; hind limb in a slightly darker brown; venter cream (Wagner et al. 2021). 
CommentIllustration: Iguana 27 (2): 10 (2014).

Behavior: diurnal, semiarboreal 
EtymologyNamed after Rike Bach (Bonn), in gratitude for supporting the second author (PG) during several fieldtrips in Indochina. 
References
  • Geißler, Peter G 2013. Eine neue Schönechse aus Indochina. Terraria-Elaphe 2013 (5): 46 - get paper here
  • Hartmann, Timo, Peter Geissler, Nikolay A. J. Poyarkov, Flora Ihlow, Eduard A. Galoyan, Dennis Rödder & Wolfgang Böhme. 2013. A new species of the genus Calotes Cuvier, 1817 (Squamata: Agamidae) from southern Vietnam. Zootaxa 3599 (3): 246–260 - get paper here
  • Karthik, Pandi; Ayuthavel Kalaimani, Rathinalingam Nagarajan 2018. An inventory on herpetofauna with emphasis on conservation from Gingee Hills, Eastern-Ghats, Southern India. Asian Journal of Conservation Biology 7 (1): 2-16 - get paper here
  • Neang T., Grismer, L.L., Hun S. & Phan C. 2015. New herpetofauna records and range extensions for Daboia siamensis (Smith, 1917) and Gekko petricolus Taylor, 1962 from Cambodia. Cambodian Journal of Natural History, 2015: 172–182 - get paper here
  • Wagner, P., Ihlow, F., Hartmann, T., Flecks, M., Schmitz, A., & Böhme, W. 2021. Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141–171 - get paper here
  • Ziegler, T & Nguyen, QT 2015. Neues von den Forschungs- und Naturschutzprojekten in Vietnam und Laos. Zeitschrift des Kölner Zoos 2015 (2): 79-
  • Ziegler, Thomas & Anna Rauhaus 2015. Erste herpetologische Amtshilfe aus Köln im Zoo Saigon. Terraria-Elaphe 2015 (4): 72-80 - get paper here
 
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