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Calotes geissleri WAGNER, IHLOW, HARTMANN, FLECKS, SCHMITZ & BÖHME, 2021

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymCalotes geissleri WAGNER, IHLOW, HARTMANN, FLECKS, SCHMITZ & BÖHME 2021: 150 
DistributionMyanmar and India (e.g., BMNH 1956.1.11.98, specimens mentioned by Lalremsanga et al. [2010]).

Type locality: Myanmar, Sagaing Division, Mon Ywa District, AK Park, Thabake Sae Camp [22.316806° N, 94.475556° E]  
Reproduction 
TypesHolotype: CAS 215539 (adult male, Fig. 6A-B, Clade B), collected by H. Win, T. Thin, S.L. Oo and H. Tun on June 9th 2000
Paratypes: CAS 210270 from Myanmar, Alaungdaw Kathapa National Park, Thabakesay (Log Cabin Camp) [22.318194° N, 94.475722° E]; ZFMK 97991 (formerly CAS 243200) from Myanmar, Chin State, Phalum District, Simggial village [23.762583° N, 93.546167° E, 1362 m.]; CAS 243028, CAS 243050 both from Myanmar, Magway Division, Gangaw District, Gangaw Township, Mauk village [22.335861° N, 94.144583° E, 205 m.]. 
DiagnosisDiagnosis: A large sized Calotes species with a known maximum SVL of 122 mm in males and 114 mm in females. Tail relatively short, up to 270 mm in males and 223 mm in females. The new species can be distinguished from other species of the complex by the combination of the following characters: 1) head and body very robust; 2) nuchal and dorsal crests continuous, composed of erect compressed scales, directed posteriorly, larger on the nuchal crest than on the dorsal crest, becoming smaller towards the tail; 3) 50–62 scale rows around midbody; 4) 35–45 vertebral spines and scales in males, 49–50 in females; 5) body scales small, homogeneous, feebly keeled and arranged in regular rows; 6) a short row of separated spines on both sides of the head, directing from the tympanum to the first scale of the nuchal crest; 7) extremities relatively short and robust; 8) oblique skin fold in front of the fore limbs, 9) head and body bluish, with a white band from the tip of the mouth along the upper lip, the tympanum and prominently continuing between the dorsolateral brownish orange body blotches on the body reaching the hind limbs, band as broad as the height of the tympanum on the head and above the front legs, becoming gradually narrower until the insertion of the hindlimbs; 10) three or more large distinct brownish orange blotches on both sides of the body between the limbs. (Wagner et al. 2021).

Color in life: Males in breeding color with blue head and body (Fig. 6C). A white band is present from the tip of the snout along the upper lips and the tympanum, predominantly continuing between the dorsolateral blotches on the lateral sides of the body to the hind limbs. The band is as broad as the height of the tympanum on the head and above the fore legs, becoming gradually narrower towards the insertion of the hind limbs. Three or more large distinct brownish-orange blotches on the lateral sides of the body between the limbs. Non-display coloration unknown. (Wagner et al. 2021). 
CommentSympatry: In India the species occurs in with Calotes jerdoni and C. versicolor (Lalremsanga et al. 2010).

Ecology. Calotes geissleri sp. n. is diurnal and semi-ar- boreal. Preferred habitats are unknown. Lalremsanga et al. (2010) collected one individual on a branch of Pi- nus kesiya, about 3m above the ground, in a secondary forest. The specimen was kept and remained greyish brown in coloration in captivity, but changed the color of the head and anterior portion of the trunk to bright blue minutes after exposure to the sun. Like other Calotes spe- cies, C. geissleri sp. n. feeds on arthropods like Coleop- tera, Formicidae and others. In India (see Lalremsanga et al. 2010) the species occurs in sympatry with Calotes jerdoni and C. versicolor (Wagner et al. 2021). 
EtymologyThe specific epithet is a patronym formed in the genitive singular honoring Dr. Peter Geißler, Museum Natur und Mensch, Freiburg, Germany, in recognition of his work on the Southeast Asian herpetofauna in general, and his collection of Calotes bachae specimens in 2009 in particular, which initiated research on the Calotes mystaceus complex. 
References
  • Lalremsanga, H.T., L. Khawlhring & Lalrotluanga 2010. Three additional lizard (Squamata: Sauria) records for Mizoram, India. Journal of Threatened Taxa 2(2): 718-720 - get paper here
  • Wagner, P., Ihlow, F., Hartmann, T., Flecks, M., Schmitz, A., & Böhme, W. 2021. Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141–171 - get paper here
 
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