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Carlia johnstonei STORR, 1974

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Higher TaxaScincidae, Eugongylinae (Eugongylini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Rough Brown Rainbow-skink 
SynonymCarlia johnstonei STORR 1974
Carlia johnstonei — COGGER 1983
Carlia johnstonei — COGGER 2000: 394
Carlia johnstonei — DOLMAN & HUGALL 2008
Carlia johnstonei — AFONSO-SILVA et al. 2017 
DistributionAustralia (Western Australia)

Type locality: Mitchell Plateau, in 14° 52’ S, 125° 50’ E, W. A. (= −14.866667, 125.833333)  
Reproductionoviparous 
TypesHolotype: WAM R43170 
DiagnosisDiagnosis: Dark blackish Carlia morphologically distinguishable from geographically overlapping species with a combination of mid-dorsal scales bicarinate (two keels), more numerous supraciliares (usually 7 vs. 6—C. amax, C. munda, C. rufilatus, C. isostriacantha sp. nov., or 5—C. gracilis), larger ear aperture with numerous sharply pointed lobules (mean of 10 lobules), but typically less than in C. insularis sp. nov. (mean of 13 lobules). Further distinguished from the latter by smaller body size (mean 36.39 mm vs. 41.83 mm), reduced head depth (mean 3.59 mm vs. 4.48 mm), shorter limbs (forelimbs 9.51 mm vs. 11.45 and hindlimbs 14.82 mm vs. 17.77 mm) and less lamellae under longest finger (mean 16.75 mm vs. 19.69 mm) and toe (mean 22.83 mm vs. 26.31 mm) [Afonso-Silva et al. 2017]. 
CommentPopulations of this species (inlcuding the paratype from East Montalivet Island, WAM R41462) have been split off as C. insularis by AFONSO-SILVA et al. 2017. 
Etymologynamed after Ronal Eric Johnstone (born 1949), Australian herpetologist. 
References
  • Afonso Silva, A. C. 2018. Evolutionary history of sympatric Rainbow skinks from the Australian Monsoonal Tropics. PhD Thesis, University of Lisbon, Portugal
  • Afonso Silva, A. C., Bragg, J. G., Potter, S., Fernandes, C., Coelho, M. M. and Moritz, C. 2017. Tropical specialist versus climate generalist: diversification and demographic history of sister species of Carlia skinks from northwestern Australia. Mol Ecol.; doi:10.1111/mec.14185 - get paper here
  • Afonso-Silva AC, Santos N, Ogilvie HA, Moritz C. 2017. Validation and description of two new north-western Australian Rainbow skinks with multispecies coalescent methods and morphology. PeerJ 5:e3724 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Couper, P., Covacevich, J., Amey, A. & Baker, A. 2006. The genera of skinks (Family Scincidae) of Australia and its island territories: diversity, distribution and identification. in: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (eds.). Evolution and Zoogeography of Australasian Vertebrates. Australian Scientific Publishing, Sydney, pp. 367-384
  • Dolman, Gaynor & Andrew F. Hugall 2008. Combined mitochondrial and nuclear data enhance resolution of a rapid radiation of Australian rainbow skinks (Scincidae: Carlia). Molecular Phylogenetics and Evolution 49 (3): 782-794 - get paper here
  • Greer, A. E. 1976. Notes on the systematics of the genus Carlia (Lacertilia: Scincidae). II. Carlia peroni (Dumeril and Bibron 1839). Herpetologica 32: 371-377 - get paper here
  • Ingram G; Covacevich J 1989. Revision of the genus Carlia (Reptilia, Scincidae) in Australia with comments on Carlia bicarinata of New Guinea. Memoirs of the Queensland Museum 27 (2): 443-490 - get paper here
  • Storr, G. M. 1974. The genus Carlia (Lacertilia: Scincidae) in Western Australia and Northern Territory. Rec. West. Aust. Mus. 3: 151-165 - get paper here
  • Storr, G. M., Smith, L. A. & Johnstone, R. E. 1981. Lizards of Western Australia. I. Skinks. Perth: University of Western Australia Press and Western Australian Museum, 200 pp.
  • Storr, G. M.; L. A. Smith, and R. E. Johnstone 1999. Lizards of Western Australia. I. Skinks. Revised Edition. Western Australian Museum
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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