You are here » home › search results › Clemmys guttata

Clemmys guttata (SCHNEIDER, 1792)

IUCN Red List - Clemmys guttata - Endangered, EN
iNaturalist logo

Can you confirm these amateur observations of Clemmys guttata?

Add your own observation of
Clemmys guttata »

Find more photos by Google images search: Google images

Higher TaxaEmydidae, Emydinae, Testudinoidea, Testudines (turtles)
Subspecies 
Common NamesE: Spotted Turtle
G: Tropfenschildkröte 
SynonymTestudo guttata SCHNEIDER 1792
Testudo anonyma SCHNEIDER 1792 (nomen nudum)
Testudo punctata SCHOEPFF 1792
Clemmys guttata — STRAUCH 1862
Emys guttata — SOWERBY 1872
Clemmys guttata — WRIGHT 1918: 56
Clemmys guttata — ERNST & BARBOUR 1989: 188
Clemmys guttata — CONANT & COLLINS 1991: 50
Clemmys guttata — CROTHER 2000
Clemmys guttata — FELDMAN & PARHAM 2002
Clemmys guttata — TTWG 2014
Chelopus guttatus — fide TTWG 2017
Melanemys guttatus — fide TTWG 2017
Geoclemys guttata — fide TTWG 2017
Nanemys guttata — fide TTWG 2017
Clemmys guttata — TTWG 2017
Clemmys guttata — TTWG 2021 
DistributionCanada (S Ontario, Quebec),
USA (along the Atlantic Coastal Plain in Maine, SE New Hampshire, Massachusetts, Connecticut, S New York, Pennsylvania, New Jersey, Delaware, Maryland, Virginia, West Virginia, North Carolina, South Carolina, Georgia, N Florida, Ohio, Indiana, Michigan, NE Illinois)

Type locality: vicinity of Philadelphia, Pennsylvania (designated by MITTLEMAN 1945)  
Reproductionoviparous. Temperature‐dependent sex determination (TSD) fide Cornejo-Páramo et al. 2020 (and references therein). 
TypesType: BMNH 1853.11.30.25 (possibly the type of Geoclemmys sebae fide P. Campbell, pers. comm., March 2019). 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2648 characters), are available for collaborators and contributors. Please contact us for details. 
CommentType species: Clemmys guttata is the type species of the genus Clemmys RITGEN, 1828. HOLMAN & FRITZ 2001 and FELDMAN & PARHAM 2002 suggested to treat Clemmys as a monotypic genus with C. guttata as its sole member. However, both author teams suggested different names for the other members of Clemmys (marmorata, muhlenbergii, insculpta).

Illustrations: Picture in Draco 13 (4): 96 (2003).

Habitat: freshwater (swamps, ponds, rivers)

Karyotype (genus Clemmys): 2n = 50: 16 macrochromosomes
with median or submedian centromeres, 10 macrochromosomes with terminal or subterminal centromeres, and 24 microchromosomes (Bickham, 1975). 
EtymologyNamed after Latin “guttata” for spotted, referring to the scattered yellow spots on the carapace.

The genus was named after Greek clemmys (κλεμμύς), tortoise. [“...Als Benennung dieser Gattung habe ich Clemmys, Diebkrotte, gewählt...”]. 
References
  • Allerstorfer, M. 2017. Die unglaublichen und faszinierenden Sinnesleistungen der Reptilien. Reptilia (Münster) 22 (125): 44-54
  • Angielczyk, Kenneth D.; Chris R. Feldman 2013. Are diminutive turtles miniaturized? The ontogeny of plastron shape in emydine turtles. Biological Journal of the Linnean Society, 2013 - get paper here
  • Anthonysamy, Whitney J. B.; Michael J. Dreslik, David Mauger, and Christopher A. Phillips 2014. A Preliminary Assessment of Habitat Partitioning in a Freshwater Turtle Community at an Isolated Preserve. Copeia Jun 2014, Vol. 2014, No. 2: 269-278. - get paper here
  • Barnwell, Mary E.;Meylan, Peter A.;Walsh, Timothy 1997. The spotted turtle (Clemmys guttata) in central Florida. Chelonian Conserv. Biol. 2 (3): 405-408
  • Beaudry, F., P.G. deMaynadier & M.L. Hunter, Jr. 2009. Seasonally Dynamic Habitat Use by Spotted (Clemmys guttata) and Blanding's Turtles (Emydoidea blandingii) in Maine Journal of Herpetology 43 (4): 636-645. - get paper here
  • Behler, John L. 1998. Wildlife as Canon sees it: Spotted Turtle (Clemmys guttata). National Geographic 193 (5): unnumbered
  • Belmore, R.E. 2004. The Spotted Turtle, Clemmys guttata. Gainesville, Florida.
  • Bentley, C.C.; Knight, J.L. 1993. The oldest spotted turtle: Clemmys guttata (Testudines: Emydidae) from the Late Pleistocene (Rancholabrean) Ardis Local Fauna, Dorchester County, South Carolina. South Carolina Geology 36:59-63
  • Bickham, John W.;Lamb, Trip;Minx, Patrick;Patton, John C. 1996. Molecular systematics of the genus Clemmys and the intergeneric relationships of Emydid turtles. Herpetologica 52 (1): 89-97 - get paper here
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598 - get paper here
  • Brown, Grover; Jonathan D. Mays, Houston C. Chandler, Benjamin S. Stegenga, Brian Kreiser & Dirk J. Stevenson 2023. Spotted Turtle (Clemmys guttata) Population Genetics in the Southeastern United States. Chelonian Conservation and Biology Dec 2023 Vol. 22, No. 2: 127-136 - get paper here
  • Buchanan, Scott W.; Bill Buffum, and Nancy E. Karraker 2017. Responses of a Spotted Turtle (Clemmys guttata) Population to Creation of Early-successional Habitat. Herpetological Conservation and Biology 12 (3) - get paper here
  • Burke, Russell L.;Leuteritz, Thomas E.;Wolfe, Alan J. 1996. Phylogenetic relationships of Emydine turtles. Herpetologica 52 (4): 572-584 - get paper here
  • Bury, R. Bruce; Ernst, Carl H. 1977. Clemmys. Catalogue of American Amphibians and Reptiles (203): 1-2 - get paper here
  • Camper, Jeffrey D. 2019. The Reptiles of South Carolina. University of South Carolina Press, 288 pp. [review in Copeia 107 (3): 590, 2019] - get paper here
  • Chandler, Houston C.; Benjamin S. Stegenga, Dirk J. Stevenson 2020. Thermal Ecology of Spotted Turtles (Clemmys guttata) in Two Southern Populations. Copeia 108 (4): 737-745 - get paper here
  • Conant, Roger 1938. The Reptiles of Ohio. American Midland Naturalist 20 (1): 1-200 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cordero, Gerardo A. 2017. Quick guide: The turtle’s shell. Biology 27: R163–R171 - get paper here
  • Cornejo-Páramo, P., Lira-Noriega, A., Ramírez-Suástegui, C. et al. 2020. Sex determination systems in reptiles are related to ambient temperature but not to the level of climatic fluctuation. BMC Evol Biol 20: 103 - get paper here
  • Cross, M.D. & Bekker, K.A. 2018. Clemmys guttata (Spotted Turtle) Fire scar healing. Herpetological Review 49 (1): 106. - get paper here
  • Cross, Matthew D. and Bekker, Kent 2017. Clemmys guttata (Spotted Turtle) Fire scarring. Herpetological Review 48 (1): 175 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • DELAY, STEPHANIE J. & JACQUELINE D. LITZGUS. 2022. CLEMMYS GUTTATA (Spotted Turtle). NEST PREDATION. Herpetological Review 53 (3): 484–485.
  • Deuss, F. 1994. Haltung und Fortpflanzung von Clemmys guttata im Zimmerterrarium. Herpetofauna 16 (92): 22-26 - get paper here
  • Ernst, C.H. 1967. A mating aggregation of the turtle Clemmys guttata Copeia 1967 (2): 473-474. - get paper here
  • Ernst, C.H. 1968. Homing ability in the spotted turtle, Clemmys guttata (Schneider) Herpetologica 24 (1): 77-78. - get paper here
  • Ernst, C.H. 1970. Reproduction in Clemmys guttata Herpetologica 26 (2): 228-232. - get paper here
  • Ernst, C.H. 1970. Home range of the spotted turtle, Clemmys guttata (Schneider) Copeia 1970 (2): 391-393. - get paper here
  • Ernst, C.H. 1975. Growth of the Spotted Turtle, Clemmys guttata Journal of Herpetology 9 (3): 313-318. - get paper here
  • Ernst, C.H. 1976. Ecology of the Spotted Turtle, Clemmys guttata (Reptilia, Testudines, Testudinidae), in Southeastern Pennsylvania Journal of Herpetology 10 (1): 25-33. - get paper here
  • Ernst, C.H. 1982. Environmental Temperatures and Activities in Wild Spotted Turtles, Clemmys guttata Journal of Herpetology 16 (2): 112-120. - get paper here
  • Ernst, C.H. & G.R. Zug 1994. Observations on the Reproductive Biology of the Spotted Turtle, Clemmys guttata, in Southeastern Pennsylvania Journal of Herpetology 28 (1): 99-102. - get paper here
  • Ernst, Carl H. 1972. Clemmys guttata. Catalogue of American Amphibians and Reptiles (124): 1-2 - get paper here
  • Ernst, Carl H. 2001. An overview of the North American turtle genus Clemmys Ritgen, 1828. Chelonian Conserv. Biol. 4 (1): 211-216
  • Ernst, Carl H.; Lovich, Jeffrey E. 2009. Turtles of the United States and Canada, 2nd edition. Johns Hopkins University Press, 827 pp. - get paper here
  • Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
  • Feldman, C. R. & Parham, James Ford 2002. Molecular Phylogenetics of Emydine Turtles: Taxonomic Revision and the Evolution of Shell Kinesis. Molecular Phylogenetics and Evolution 22 (3): 388-398 - get paper here
  • Feng, Christina Y.; David Mauger, Jason P. Ross, and Michael J. Dreslik 2019. Size and Structure of Two Populations of Spotted Turtle (Clemmys guttata) at Its Western Range Limit. Herp. Conserv. Biology 14 (3): - get paper here
  • Folkerts, G.W. & A.C. Skorepa 1967. A Spotted Turtle, Clemmys guttata (Schneider), from Southeastern Georgia Herpetologica 23 (1): 63. - get paper here
  • Froom, B. 1976. The turtles of Canada. McClelland & Stewart, Toronto 120 pp. - get paper here
  • Graham, Terry E. 1970. Growth Rate of the Spotted Turtle, Clemmys guttata, in Southern Rhode Island. Journal of Herpetology 4 (1-2): 87-88 - get paper here
  • Gray, Brian S. and Paul Curtis 2003. A note on sexual dichromatism in Clemmys guttata from Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 38(6):120-121 - get paper here
  • Graziano, M; 2019. Clemmys guttata (Spotted Turtle) Male combat. Herpetological Review 50 (2): 349-350 - get paper here
  • Green, N.B., & Pauley, T.K. 1987. Amphibians and reptiles in West Virginia. Univ. of Pittsburgh Press, Pittsburgh, 241 pp.
  • HARTZELL, SEAN M. 2022. Geographic distribution: CLEMMYS GUTTATA (Spotted Turtle). USA: PENNSYLVANIA: Columbia Co. Herpetological Review 53 (1): 76.
  • Haxton, T. & M. Berrill 2001. Seasonal Activity of Spotted Turtles (Clemmys guttata) at the Northern Limit of Their Range Journal of Herpetology 35 (4): 606-614. - get paper here
  • Holman, J. Alan; Fritz, Uwe 2001. A new emydine species from the Middle Miocene (Barstovian) of Nebraska, USA with a new generic arrangement for the species of Clemmys sensu McDOWELL (1964). Zoologische Abhandlungen<br>(Dresden) 51(2):331-353 - get paper here
  • Howell, Hunter J. and Richard A. Seigel 2018. An Examination of the Accuracy of Using Plastral Scute Rings to Age Spotted Turtles (Clemmys guttata). Chelonian Conservation and Biology Jun 2018, Vol. 17, No. 1: 104-108. - get paper here
  • Howell, Hunter J. and Richard A. Seigel 2019. The Effects of Road Mortality on Small, Isolated Turtle Populations. Journal of Herpetology 53 (1): 39 - get paper here
  • Howell, Hunter J., Donald G. McKnight and Richard A. Seigel. 2016. A novel method of collecting Spotted Turtles (Clemmys guttata). Herpetological Review 47 (2): 202-205 - get paper here
  • Howell, Hunter J.; Richard H. Legere, David S. Holland & Richard A. Seigel 2019. Long-Term Turtle Declines: Protected Is a Verb, Not an Outcome. Copeia 107 (3): 493-501 - get paper here
  • Hunter, M.L., Jr., Albright, J., & Arbuckle, J., eds. 1992. The amphibians and reptiles of Maine. Maine Agric. Experiment Station, Orono, Bull. 838 188 pp.
  • Jamniczky, H. A. and A. P. Russell 2004. A geometric morphometric assessment of the 'batagurine process' of testudinoid turtles. Amphibia-Reptilia 25 (4): 369-379 - get paper here
  • Jensen, John B.; Carlos D. Camp, Whit Gibbons, & Matt J. Elliott 2008. Amphibians and reptiles of Georgia. University of Georgia Press, 575 pp.
  • King, Richard B.; Oldham, Michael J.; Weller, Wayne F.; Wynn, Douglas. 1997. Historic and current amphibian and reptile distributions in the Island Region of Western Lake Erie. American Midland Naturalist 138(1):153-173 - get paper here
  • Krolak, Monika and Hans-Jürgen Bidmon. 2016. Klärung von Gesundheitsproblemen im Internet: Hilfegesuch bei der Haltung einer Tropfenschilkröte Clemmys guttata. Schildkröten im Fokus 13 (2): 22-25. [hjb@hirn.uni-duesseldorf.de]
  • Krolak, Monika and Rolf Krolak. 2015. Schwarzgoldene Juwelen im Garten: Ganzjährige geschützte Freilandgruppenhaltung der Tropfenschildkröte, Clemmys guttata. Schildkröten im Fokus. 12 (1): 3-18
  • Lambertz, H. & Lambertz, K. 2002. Langjährige Zuchterfolge von Clemmys guttata. Elaphe 10 (1): 37-41
  • Litzgus, J.D. 2006. Sex Differences in Longevity in the Spotted Turtle (Clemmys guttata) Copeia 2006 (2): 281-288. - get paper here
  • Litzgus, J.D. & R.J. Brooks 1998. Reproduction in a Northern Population of Clemmys guttata Journal of Herpetology 32 (2): 252-259. - get paper here
  • Litzgus, J.D. & R.J. Brooks 2000. Habitat and Temperature Selection of Clemmys guttata in a Northern Population Journal of Herpetology 34 (2): 178-185. - get paper here
  • Litzgus, J.D. & T.A. Mousseau 2003. Multiple Clutching in Southern Spotted Turtles, Clemmys guttata Journal of Herpetology 37 (1): 17-23. - get paper here
  • Litzgus, J.D. & T.A. Mousseau 2004. Home range and seasonal activity of Southern Spotted Turtles (Clemmys guttata): Implications for management Copeia 2004 (4): 804-817. - get paper here
  • Litzgus, Jacqueline D. & Timothy A. Mousseau 2006. Geographic variation in reproduction in a freshwater turtle (Clemmys guttata). Herpetologica 62 (2): 132-140 - get paper here
  • Litzgus, Jacqueline D., Frances Bolton and Albrecht I. Schulte-Hostedde. 2008. Reproductive output depends on body condition in spotted turtles, Clemmys guttata. Copeia 2008 (1): 86-92 - get paper here
  • Litzgus, Lacqueline D.;Brooks, Ronald J. 1998. Testing the validity of counts of plastral scute rings in spotted turtles, Clemmys guttata. Copeia 1998 (1): 222-225 - get paper here
  • Lovich, J.E. 1988. Geographic Variation in the Seasonal Activity Cycle of Spotted Turtles, Clemmys guttata Journal of Herpetology 22 (4): 482-485. - get paper here
  • Lovich, J.E. & Jaworski, T.R. 1988. Annotated checklist of Amphibians and Reptiles reported from Cedar Bog, Ohio. Ohio J. Sci. 88 (4): 139-143 - get paper here
  • Lovich, Jeffrey E.; Carl H. Ernst, Evelyn M. Ernst, and Julia L. Riley 2014. A 21-Year Study of Seasonal and Interspecific Variation of Hatchling Emergence in a Nearctic Freshwater Turtle Community: To Overwinter or Not To Overwinter? Herpetological Monographs, 28: 93-109 - get paper here
  • Manns, B. 1969. Notes on Clemmys guttata as a scavenger in a Parkville, Maryland pond Bull. Maryland Herpetol. Soc., 5(2): 57. - get paper here
  • McCord, W.P. & Joseph-Ouni, M. 2006. Chelonian Illustrations # 24 - North American Wood and Pond Turtles. Reptilia (GB) (44): 83-86 - get paper here
  • Mehrtens, J.M. 1949. Copulation of Clemmys guttata and Pseudemys troostii elegans Herpetologica 5 (6): 150. - get paper here
  • Merkle, D. A. 1975. A taxonomic analysis of the Clemmys complex (Reptilia: Testudines) utilizing starch gel electrophoresis. Herpetologica 31 (2): 162-166 - get paper here
  • Milam, J.C. & S.M. Melvin 2001. Density, Habitat Use, Movements, and Conservation of Spotted Turtles (Clemmys guttata) in Massachusetts Journal of Herpetology 35 (3): 418-427. - get paper here
  • Mitchell, J. C. & Reay, K.K. 1999. Atlas of amphibians and reptiles in Virginia. Specialty Publication 1, VA Dept of Game and Fisheries, 122 pp. - get paper here
  • Mitchell, J.C. 1994. The reptiles of Virginia. Virginia Department of Game and Inland Fisheries, ca. 350 pp.
  • MORRISON, BENJAMIN R.; STEVEN G. PLATT & THOMAS R. RAINWATER. 2022. CLEMMYS GUTTATA (Spotted Turtle). ATTEMPTED PREDATION. Herpetological Review 53 (2): 309–310.
  • Moski, H.C. 1957. Algal occurrence on the turtle Clemmys guttata Copeia 1957 (1): 50-51. - get paper here
  • MOTA, JOEL L.; JONATHAN DRESCHER-LEHMAN, DONALD J. BROWN, THOMAS S. AKRE & KEVIN J. OXENRIDER. 2022. Geographic distribution: CLEMMYS GUTTATA (Spotted Turtle). USA: WEST VIRGINIA: Hardy Co. Herpetological Review 53 (2): 260–261.
  • Netting, M. 1936. Hibernation and Migration of the Spotted Turtle, Clemmys guttata (Schneider). Copeia, 1936(2), 112-112 - get paper here
  • Netting, M. Graham 1940. The spotted turtle, Clemmys guttata (Schneider) (an addition to the herpetofauna of West Virginia). Proceedings, West Virginia Academy of Science 14: 146-147
  • Niederberger, Allan J.;Seidel, Michael E. 1997. Geographic Distribution. Clemmys guttata. Herpetological Review 28 (3): 156 - get paper here
  • Oxenrider, K J; Heres, B M & Brown, D J 2018. Clemmys guttata (Spotted Turtle) Habitat use. Herpetological Review 49 (3): 525-526 - get paper here
  • Oxenrider, K J; Heres, B M; Mota, J L & Brown, D J 2019. Influence of Bait Type on Capture Success of Clemmys guttata and Chrysemys picta Using Small Hoop Nets in Shallow Wetlands. Herpetological Review 50 (3): 490-492 - get paper here
  • Palmer, W.M. & Braswell, A.L. 1995. Reptiles of North Carolina. Univ. North Carolina Press
  • Parker, Patricia G.;Whiteman, Howard H. 1993. Genetic diversity in fragmented populations of Clemmys guttata and Chrysemys picta marginata as shown by DNA fingerprinting. Copeia 1993 (3): 841-846 - get paper here
  • Paterson, J. E., Pulfer, T., Horrigan, E., Sukumar, S., Vezina, B. I., Zimmerling, R., & Davy, C. M. 2021. Individual and synergistic effects of habitat loss and roads on reptile occupancy. Global Ecology and Conservation, 31, e01865 - get paper here
  • Perry, David A. 2018. Herpetological Survey of Ware Creek Wildlife Management Area. Catesbeiana 38 (2): 93-103 - get paper here
  • Phillips, JG 2016. Updated Geographic Distributions of Michigan Herpetofauna: a Synthesis of Old and New Sources. The Journal of North American Herpetology 2016: 45–69 - get paper here
  • POGUE, CONNOR D. & BLAISE G. BYRD. 2022. Geographic distribution: CLEMMYS GUTTATA (Spotted Turtle). USA: NORTH CAROLINA: Greene Co. Herpetological Review 53 (1): 75–76.
  • Rasmussen, M.L. & J.D. Litzgus 2010. Habitat Selection and Movement Patterns of Spotted Turtles (Clemmys guttata): Effects of Spatial and Temporal Scales of Analyses Copeia 2010 (1): 86-96. - get paper here
  • Ritgen, F.A. 1828. Versuch einer natürlichen Eintheilung der Vögel. Nova Acta physico-medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum, 14: 245–284.
  • Roe, John H. and Nacy, Joseph P. 2017. Terrapene carolina carolina (Eastern Box Turtle) and Clemmys guttata (Spotted Turtle) Interspecific interaction. Herpetological Review 48 (1): 181-182 - get paper here
  • Rowe, J. W., J. R. Gradel, et al. 2012. Sexual dimorphism in size and shell shape, and dichromatism of Spotted Turtles (Clemmys guttata) in southwestern Michigan. Amphibia-Reptilia 33(3-4): 443-450 - get paper here
  • Rowe, John W.; Jessica R. Gradel, and Charles F. Bunce 2012. Effects of Weather Conditions and Drought on Activity of Spotted Turtles (Clemmys guttata) in a Southwestern Michigan Wetland. The American Midland Naturalist Jan 2013, Vol. 169, No. 1: 97-110. - get paper here
  • Ruther, E., Degregorio, B., Sperry, J. & Sekscienski, S. 2017. Clemmys guttata (Spotted Turtle) Longevity. Herpetological Review 48 (4): 834 - get paper here
  • Schaefer, I. 1994. Haltung und Zucht der Tropfenschildkröte (Clemmys guttata). Salamandra 30 (1): 22-32 - get paper here
  • Schneider, J. G. 1792. Beschreibung und Abbildung einer neuen Art von Wasserschildkröte nebst Bestimmungen einiger bisher wenig bekannten fremden Arten. Schriften der Gesellschaft Naturforschender Freunde zu Berlin 10: 259-283 [1791] - get paper here
  • Seburn, David C. 2012. Why didn’t the Spotted Turtle (Clemmys guttata) cross the road? Herpetology Notes 6: 527-530
  • Seidel, Michael E. & Carl H. Ernst 2017. A Systematic Review of the Turtle Family Emydidae. Vertebrate Zoology 67 (1): 1–122 - get paper here
  • Sowerby, James de Carle and Edward Lear 1872. Tortoises, terrapins, and turtles: drawn from life. London, Paris, and Frankfort :H. Sotheran, J. Baer & co. - get paper here
  • Spinks, Phillip Q. & H. Bradley Shaffer 2009. CONFLICTING MITOCHONDRIAL AND NUCLEAR PHYLOGENIES FOR THE WIDELY DISJUNCT EMYS (TESTUDINES: EMYDIDAE) SPECIES COMPLEX, AND WHAT THEY TELL US ABOUT BIOGEOGRAPHY AND HYBRIDIZATION. Systematic Biology 58(1): 1-20 - get paper here
  • Spinner, L. 2010. Gefleckte Spaβmacher - Die Tropfenschildkröten (Clemmys guttata) der Cape-Cod-Halbinsel Terraria (24) 5 (4): 60-67.
  • Steen, David A. 2013. Clemmys guttata (spotted turtle) reproduction. Herpetological Review 44 (3): 501 - get paper here
  • Steiner, H. M. 1977. Lebensraumwahl seniler Clemmys c. caspica in der Osttürkei (Reptilia, Testudines, Emydidae). Salamandra 13 (1): 53 - get paper here
  • Stevenson, D J; Mays, J D & Chandler, H C 2018. Clemmys guttata (Spotted Turtle) Coloration. Herpetological Review 49 (2): 317-318 - get paper here
  • STEVENSON, DIRK J. et al. 2021. New County Records for Amphibians and Reptiles in Georgia, USA. Herpetological Review 52 (2): 350-359
  • TTWG [Peter Paul van Dijk, John B. Iverson, Anders G.J. Rhodin, H. Bradley Shaffer, and Roger Bour] 2014. Turtles of the World, 7th Edition: Annotated Checklist of Taxonomy, Synonymy, Distribution with Maps, and Conservation Status. 000.v7. Chelonian Research Monographs (ISSN 1088-7105) No. 5, doi:10.3854/crm.5.000.checklist.v7.2014 - get paper here
  • TTWG; Anders G.J. Rhodin, John B. iverson, Roger Bour, Uwe Fritz, Arthur GeoRges, H. BRadley Shaffer, and Peter Paul van Dijk 2017. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (8th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Pritchard, P.C.H., and Mittermeier, R.A. (Ed Chelonian Research Monographs 7:1–292. doi: 10.3854/crm.7.checklist.atlas.v8.2017 - get paper here
  • TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
  • ULTSCH, G. R., AND D. M. CARROLL 2020. Climate Change and the Timing of Emergence from Hibernation of Spotted Turtles (Clemmys guttata) in Southeastern New Hampshire. Herpetological Review 51: 18-20.
  • Walker, C. 1931. Notes on Reptiles in the Collection of the Ohio State Museum. Copeia 1931 (1): 9-13 - get paper here
  • Ward, F. P.;Hohmann, C. J.;Ulrich, J. F.;Hill, S. E. 1976. Seasonal microhabitat selections of spotted turtles (Clemmys guttata) in Maryland elucidated by radioisotope tracking. Herpetologica 32 (1): 60-64 - get paper here
  • WEBER, ANDREW S. 2022. CLEMMYS GUTTATA (Spotted Turtle). USE OF ANTHROPOGENIC MATERIAL. Herpetological Review 53 (3): 485–486.
  • Wilson, D.S. et al. 2003. Estimating age of turtles from growth rings: a critical evaluation of the technique. Herpetologica 59 (2): 178-194 - get paper here
  • Wilson, T.P. 2000. Notes on Eggshell Type of the Spotted Turtle, Clemmys guttata: Geographic Variation or Differences in Definition? Bull. Maryland Herpetol. Soc., 36(2): 68-69. - get paper here
  • Wright, A.H. 1918. Notes on Clemmys. Proc. Biol. Soc. Washington 31: 51-58 - get paper here
  • Yagi, Katharine T. and Jacqueline D. Litzgus 2012. The Effects of Flooding on the Spatial Ecology of Spotted Turtles (Clemmys guttata) in a Partially Mined Peatland. Copeia 2012 (2): 179-190. - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Clemmys&species=guttata

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator