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Cnemaspis muria RIYANTO, MUNIR, MARTAMENGGALA, FITRIANA & HAMIDY, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Muria Rock Gecko
Indonesian: Cicak Batu Gunung Muria 
SynonymCnemaspis muria RIYANTO, MUNIR, MARTAMENGGALA, FITRIANA & HAMIDY 2019 
DistributionIndonesia (Java)

Type locality: river bank at Gunung Muria, Kajar (village), Dawe (District), Kudus (Regency), Jawa Tengah (Province), Indonesia (06o39’47.4” S; 110o53’22.9” E; elevation 599 m asl  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype. MZB Lace. 14571 (Fig. 2A), an adult male, collected on 11 August 2018 by Awal Riyanto, Misbahul Munir, Rubby Alfian, Lianita Rarasandy and Rega D. Ganiarta.
Paratypes. MZB. Lace. 14564 (Fig. 2B), an adult female; MZB. Lace. 14565–70, six adult males from the same locality as holotype (06 ̊39’33.5” S; 110 ̊53’20.6” E; elevation 650 m asl), collected on 17 July 2018 by Andri I.S. Martamenggala, and MZB. Lace.14572 (Fig. 2C), an adult male, with the same data as holotype (6°39’36.9” S; 110o53’20.0” E; elevation 646 m asl). 
DiagnosisDiagnosis: Cnemaspis muria sp. nov. differs from its congeners in Southeast Asian by the following combination characters: (1) maximum SVL of at least 58.1 mm in males and 56.9 mm in females, (2) a pair of sharpe conical tubercle clusters on occiput, (3) nuchal loop bearing a bridge of warts from the upper tympanum to the nape and made in a curved line, (4) dorsal tubercles not linearly arranged, (5) 18–20 paravertebral tubercles, (6) postmental separated by a single scale, (7) gular, pectoral, abdominal, subantebrachial, subbrachial, subfemoral, subtibial and subcaudal scales keeled, (8) no tubercles on lower flank, (9) both precloacal and femoral pores absent, (10) enlarged submetacarpal scales present at the base of first finger, (11) enlarged submetatarsal present at the base of first toe , (12) 38–40 ventral scales, (13) 31–35 lamellae under fourth toe, (14) two postcloacal tubercles on each side, (15) enlarged median subcaudal scale row present, (16) caudal tubercles encircling tail, (17) sexually dimorphic in color pattern: males with a yellow belly and the proximal subcaudal surfaces yellow becoming white distally, female with a white belly with proximal subcaudal surface of alternating white and black rings and black coloration distally.


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CommentHabitat: Cnemaspis muria sp. nov. is a scansorial species known only from large granite rock micro- habitats along rivers and coffee plantations (Fig. 8) on the southern slope of Gunung Muria at middle elevations, between 560 and 599 m. The holotype was caught at night, hanging on a tree root, 2 m above a dry river bank. The paratypes MZB. Lace. 14564 and MZB. Lace. 14572 were also caught at night, whereas the other paratypes were caught during the day hanging on shaded crevices of rocks. MZB. Lace. 14572 and four another specimens not col- lected were found foraging on rock walls, and sympatric with Cyrtodactylus sp. 
EtymologyThe specific epithet muria is a noun in apposition and refers to Gunung Muria, the type locality of this species, and so far, the only known locality for the genus Cnemaspis in Java. 
References
  • RIYANTO, AWAL; MISBAHUL MUNIR, ANDRI I. S. MARTAMENGGALA, YULI SULISTYA FITRIANA, AMIR HAMIDY 2019. Hiding in plain sight on Gunung Muria: A new species and first record of rock gecko (Cnemaspis Strauch, 1887; Squamata, Gekkonidae) from Java, Indonesia. Zootaxa 4608 (1): 155–173 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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