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Cnemaspis hangus GRISMER, WOOD, ANUAR, RIYANTO, AHMAD, MUIN, SUMONTHA, GRISMER, ONN, QUAH & PAUWELS, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCnemaspis hangus GRISMER, WOOD, ANUAR, RIYANTO, AHMAD, MUIN, SUMONTHA, GRISMER, ONN, QUAH & PAUWELS 2014: 86 
DistributionPeninsular Malaysia (Pahang)

Type locality: Bukit Hangus, Pahang, Peninsular Malaysia (04°16.142’N, 102°13.370’E) at 10 m in elevation  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: UKMHC (given as HC) 00227, Adult male collected on 24 June 2008 by Chan Kin Onn at 1030 hrs. Paratypes. Adult female (HC 00225) has the same collection data as the holotype. 
DiagnosisDiagnosis: Cnemaspis hangus sp. nov. differs from all other Southeast Asia species of Cnemaspis in having the unique combination of adult males reaching 50.5 mm SVL, adult females reaching 47.0 mm SVL; nine supralabials; eight infralabials; ventrals keeled; no precloacal pores; moderately prominent dorsal tubercles; 22–24 paravertebral tubercles; dorsal body tubercles semi-randomly arranged; tuberculation weak on flanks; caudal tubercles not encircling tail; tubercles may be within lateral caudal furrows anteriorly only; lateral row of caudal tubercles present; ventrolateral caudal tubercles absent; subcaudals keeled; no enlarged, median subcaudal scale row; two postcloacal tubercles; no enlarged femoral, subtibial, or submetatarsal scales; subtibials keeled; and 27–34 subdigital lamellae on fourth toe (Tables 6,7). Cnemaspis hangus sp. nov. lacks the diagnostic color pattern characteristics of other species in the Peninsular clade. 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet hangus is an invariable noun in apposition in reference to the Malay word “hangus” which means to burn or scorch and refers to this species’ overall burnt appearance in the dark color pattern phase. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; P. L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & NORHAYATI AHMAD 2015. A new insular species of Rock Gecko (Cnemaspis Boulenger) from Pulau Langkawi, Kedah, Peninsular Malaysia. Zootaxa 3985 (2): 203–218 - get paper here
  • Grismer, L.L.; P. L. WOOD, JR., SHAHRUL ANUAR, H. R. DAVIS, A. J. COBOS & M. L. MURDOCH 2016. A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia’s endemic karst forest herpetofauna and the need for its conservation. Zootaxa 4061 (1): 001–017 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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