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Cnemaspis niyomwanae GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Niyomwan’s Rock Gecko
Thai: Djing Djok Niew Yaow Niyomwan 
SynonymCnemaspis niyomwanae GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA 2010
Cnemaspis niyomwanae — GRISMER et al. 2014: 36 
DistributionNE Thailand (border regions of Trang and Satun provinces), W Cambodia

Type locality: Thum Khao Ting, Palean District, Trang Province, Thailand (07°09.943N 99°48.142E) at 28 m in elevation.  
Reproductionoviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: THNHM 15910, adult female. Collected by M. Sumontha on 26 December 2009. Paratypes. Adult male (ZMKU Rep-000315), adult female (PSUZC RT 2010.56), subadult males (CUMZ R-2009,6,24-10, KZM 008) from Baan Nam Pud, La-ngu District, Satun Province (07°05.688N 99° 54.732E) at 38–46 m in elevation. ZMKU Rep-000315, CUMZ R-2009,6,24-10 and KZM 008 were collected by Siriwat Dangsri on 6 and 7 October 2009. PSUZC RT 2010.56 was collected by Thanin Kaewmanee on 24 December 2009. 
DiagnosisDiagnosis: Adult males reaching 45.9 mm SVL, adult females reaching 56.8 mm SVL; 8–11 supralabials; 6–8 infralabials; gulars smooth; forearm scales keeled; subtibials, ventrals, subcaudals smooth; dorsal tubercles keeled; 26–31 paravertebral tubercles; enlarged, elongate, laterally compressed, white, isolated tubercles on sides of neck, shoulders, and flanks; dorsolateral, lateral, and ventrolateral caudal tubercles usually present only anteriorly; caudal tubercles do not encircle tail; caudal tubercles absent from lateral, caudal furrow; middorsal caudal furrow absent; median row of smooth, enlarged subcaudals; three pore- bearing, precloacal scales in males; one or two postcloacal tubercles; shield-like subtibials and enlarged, submetatarsals absent; 31–34 subdigital lamellae on fourth toe; yellow and orange alternating bands on forearms and forelimbs in subadult and adult males; paired, yellow, rostral stripes in subadult and adult males; paried, cream to white, circular, occipital blotches; large, cream to white, paravertebral butterfly-shaped dorsal blotches; no dark, longitudinal, gular markings or blotches; head not yellow in adult males; no black neck patch enclosing a white to yellow ocellus; no dark shoulder patch enclosing white to yellow ocellus; no prominent, yellow to white, prescapular crescent or transverse bars on flanks. These differences are summarized across all species in TABLES 1 and 2 in GRISMER et al. 2010.


Additional details (951 characters) are available for collaborators and contributors. Please contact us for details. 
CommentDistribution: See map in Grismer et al. 2014: 18 (Fig. 4). 
EtymologyThis species is named in honor of Ms. Piyawan Niyomwan, Thai herpetologist who has worked for many years making significant contributions to our knowledge of the distribution of the amphibians and reptiles of Thailand. 
References
  • Ampai N, Wood Jr PL, Stuart BL, Aowphol A 2020. Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159 - get paper here
  • GRISMER, L. LEE; MONTRI SUMONTHA, MICHAEL COTA, JESSE L. GRISMER, PERRY L. WOOD, JR., OLIVIER S. G. PAUWELS & KIRATI KUNYA 2010. A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa 2576: 1–55 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • QUAH, EVAN S. H.; PERRY L. JR. WOOD, M.S. SHAHRUL ANUAR, MOHD ABDUL MUIN 2020. A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia with an updated checklist of the herpetofauna of Tuba Island. Zootaxa 4767 (1): 138–160 - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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