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Cnemaspis heteropholis BAUER, 2002

IUCN Red List - Cnemaspis heteropholis - Near Threatened, NT

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCnemaspis heteropholis BAUER 2002: 161
Cnemaspis heteropholis — MANAMENDRA-ARACHCHI et al. 2007
Cnemaspis heteropholis — KHANDEKAR et al. 2020 
DistributionIndia (Gund, Uttar Kannada, Goa)

Type locality: North Kanara, Karnataka State; Gund (15.167° N, 74.667° E), Uttar Kannada Dis- trict, Karnataka, India, fide Khandekar et al. 2020.  
Reproductionoviparous 
TypesHolotype: ZMH R06158, adult female, collected by G. A. Von Maydell on 20 January 1956. 
DiagnosisDiagnosis: A medium-sized Cnemaspis, snout-vent length upto 45 mm; dorsal scales on trunk heterogeneous, small, weakly keeled, oval, granular scales intermixed with enlarged, irregularly arranged, strongly keeled, conical tubercles; tubercles on dorsolateral aspect of flank more pronounced and conical than those on paravertebral region; spine-like scales absent on flanks. Ventral scales on belly smooth, subimbricate, 20–25 scales across mid-body, 119–123 longitudinal scales between mental to anterior border of cloaca. Subdigital scansors smooth, entire except for three or four proximal and 1–3 distal scansors on digit I–V of both manus and pes which are divided, unnotched; 20–22 lamellae under digit IV of manus, and 24 or 25 lamellae under digit IV of pes. Males with five or six femoral pores on each thigh, separated by 16–18 poreless scales; precloacal pores absent; dorsal pholidosis of tail homogeneous, composed of small, smooth, flattened, roughly circular, regularly arranged, subimbricate scales; without whorls of enlarge tubercles; median row of sub-caudals smooth, enlarged; enlarged median row is irregularly arranged with a large scale alternating with two slightly smaller scales on the posterior portion of the tail; the larger scales on the anterior portion divided; postcloacal spur small, indistinct, not conical (Khandekar e tal. 2020).


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CommentKhandekar et al. 2020 restricted Cnemaspis heteropholis to the vicinity of its type locality and expand the morphological description of the species by including first confirmed male from its type locality and one male and two females from additional localities. They then describe the Sakleshpur population as a new species, Cnemaspis magnifica.

Ganesh et al. (2011) provided the first description of a male C. heteropholis from Agumbe (13°50’N, 75°09’E; 557 m asl.), Shimoga district, Karnataka, ca. 200 km south from its type locality. However, Khandekar et al. 2020 state that “the species reported as C. heteropholis by Ganesh at al. (2011) from Agumbe is in fact yet another undescribed species (Pal et al. in prep.).” 
EtymologyNamed after the Greek “heteros” = different and “pholis” = scale, in reference to the strongly heterogenous dorsal scalation typical of this species. 
References
  • Bauer, A.M. 2002. Two new species of Cnemaspis (Reptilia: Squamata: Gekkonidae) from Gund, Uttara Kannada, India. Mitt. Hamburg. Zool. Mus. Inst. 99: 155-167
  • Biswas, Sayantan 2008. A Possible Occurrence of Regional Integumentary Loss in Cnemaspis heteropholis from Southern India. Gekko 5 (2): 28-30
  • Ganesh, S.R., Sreekar, R., Pal, S.P., Ramchandra, G., Srinivasulu, C. & Srinivasulu, B. 2011. Discovery and first description of male Cnemaspis heteropholis Bauer, 2002 (Reptilia: Gekkonidae) from Agumbe, central Western Ghats, India. Journal of Threatened Taxa, 3, 2023–2027 - get paper here
  • KHANDEKAR, AKSHAY; TEJAS THACKERAY, SAUNAK PAL & ISHAN AGARWAL 2020. A new large-bodied, rupicolous Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) allied to Cnemaspis heteropholis Bauer, 2002 from the Central Western Ghats of Karnataka, India. Zootaxa 4801 (1): 057–084 - get paper here
  • Manamendra-Arachchi, Kelum; Batuwita, Sudesh & Pethiyagoda, Rohan 2007. A taxonomic revision of the Sri Lankan day-geckos (Reptilia: Gekkonidae: Cnemaspis), with description of new species from Sri Lanka and southern India. Zeylanica 7 (1): 9-122
  • Sayyed A, Pyron RA, Dileepkumar R. 2018. Four new species of the genus Cnemaspis Strauch, (Sauria: Gekkonidae) from the northern Western Ghats, India. Amphibian & Reptile Conservation 12(2) [General Section]: 1–29 (e157) - get paper here
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
 
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