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Cnemaspis adangrawi AMPAI, RUJIRAWAN, WOOD, STUART & AOWPHOL, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Adang-Rawi Rock Gecko
Thai: Jing Jok Niew Yaow Ko Adang-Rawi 
SynonymCnemaspis adangrawi AMPAI, RUJIRAWAN, WOOD, STUART & AOWPHOL 2019 
DistributionThailand (Satun)

Type locality: Thailand, Satun Province, Mueang Satun District, Tarutao National Park, Adang Island, Jonsalad Waterfall (6°30.7806'N, 99°18.0072'E; 84 m a.s.l.; Fig. 13A in Ampai et al. 2019.  
Reproduction 
TypesHolotype: ZMKU R 00767, adult male, collected on 9 November 2017 by Natee Ampai, Attapol Rujirawan, Siriporn Yodthong, and Korkwan Termprayoon.
Paratypes. Fourteen paratypes (adult males = 10, adult females = 4). ZMKU R 00768 (1 adult female), same locality and collectors as holotype. ZMKU R 00771 (1 adult female), and ZMKU R 00769–00770, THNHM 28206–28209 (6 adult males), same data as holotype except collected 6 April 2018. ZMKU R 00773, ZMKU R 00775, THNHM 28210 (3 adult males) and ZMKU R 00774 (1 adult female), same collectors as holotype except from Rawi Island (6°33.9084'N, 99°15.5088'E; 7 m a.s.l.; Fig. 13B), collected on 7 April 2018. ZMKU R 00776 (1 adult male) and THNHM 28211 (1 adult female), same collectors as holotype except from Rawi Island (6°33.3474'N, 99°15.0018'E; 7 m a.s.l.; Fig. 13C), collected on 8 April 2018. 
DiagnosisDiagnosis. Cnemaspis adangrawi sp. nov. can be distinguished from all other Cnemaspis by having the following combination of characters: (1) adult males with maximum SVL length 44.9 mm (mean 41.8 ± SD 2.5, n = 11) and females with maximum SVL 43.8 mm (mean 42.5 ± SD 1.5, n = 4); (2) 10 supralabials and 9 infralabials; (3) 6–8 pore-bearing precloacal scales with rounded pores arranged in chevron shape and separated; (4) 23–25 paravertebral tubercles randomly arranged; (5) 26–28 subdigital lamellae under the 4th toe; (6) subcaudal scales keeled and lacking enlarge median row; (7) one postcloacal tubercle each side; (8) gular region, abdomen, limbs and subcaudal region yellowish in males only; (9) mid-gular marking absent in males and females. These differences are summarized for geographically close congeners in the siamensis group (Table 6 in Ampai et al. 2019).

Comparisons. Cnemaspis adangrawi sp. nov. can be distinguished from other members of the siamensis group (C. chanardi, C. huaseesom, C. omari, C. phangngaensis, C. punctatonuchalis, C. roticanai, C. siamensis, C. thachanaensis, and C. vandeventeri) by having a smaller maximum SVL of 44.9 mm (vs 47.0 mm in C. roticanai, 49.6 mm in C. punctatonuchalis) and by having a larger maximum SVL 44.9 mm (vs 40.9 mm in C. chanardi, 43.5 mm in C. huaseesom, 41.3 mm in C. omari, 42.0 mm in C. phangngaensis, 39.7 mm in C. siamensis, 39.0 mm in C. thachanaensis, and 44.7 mm in C. vandeventeri).
Cnemaspis adangrawi sp. nov. is distinguished from C. omari, C. punctatonuchalis, C. roticanai, C. siamensis, and C. vandeventeri by having 10 supralabial scales (vs eight in C. punctatonuchalis and 8–9 in C. omari, C. roticanai, C. siamensis, and C. vandeventeri). This species is distinguished from C. chanardi, C. omari, C. phangngaensis, C. punctatonuchalis, C. roticanai, and C. siamensis by having 9 infralabial scales (vs 8 in C. chanardi, 7–8 in C. omari, C. punctatonuchalis, C. roticanai, 10 in C. phangngaensis, and 6–8 in C. siamensis). This species is distinguished from C. huaseesom and C. punctatonuchalis by having keeled ventral and subcaudal scales (vs smooth ventral and subcaudal scales in C. huaseesom and C. punctatonuchalis).
Cnemaspis adangrawi sp. nov. is distinguished from C. phangngaensis and C. vandeventeri by having 6–8 precloacal pores (vs 4 in C. phangngaensis and C. vandeventeri). This species is distinguished from C. punctatonuchalis, C. siamensis, and C. thachanaensis by presence of precloacal pores (vs precloacal pores absent in C. punctatonuchalis, C. siamensis, and C. thachanaensis). This species is distinguished from C. huaseesom and C. phangngaensis by having a separated row of precloacal pores (vs continuous in C. huaseesom and C. phangngaensis).
Cnemaspis adangrawi sp. nov. is distinguished from C. phangngaensis and C. thachanaensis by having 23–25 paravertebral tubercles (vs 22 in C. phangngaensis and 15–19 in C. thachanaensis). This species is distinguished from C. huaseesom, C. omari, C. punctatonuchalis, C. roticanai, C. siamensis, C. thachanaensis, and C. vandeventeri by lacking tubercles on lower flanks (vs present in C. huaseesom, C. omari, C. punctatonuchalis, C. roticanai, C. siamensis, C. thachanaensis, and C. vandeventeri). This species is distinguished from C. phangngaensis, C. punctatonuchalis, and C. thachanaensis by having 26–28 lamellae under 4th toe (vs 29 in C. phangngaensis, 29–31 in C. punctatonuchalis, and 24 in C. thachanaensis). This species is distinguished from C. chanardi and C. vandeventeri by having lateral caudal furrows (vs lacking in C. chanardi and C. vandeventeri).
Cnemaspis adangrawi sp. nov. can be further distinguished from C. chanardi, C. punctatonuchalis, C. siamensis, and C. vandeventeri by lacking enlarged median subcaudal scales (vs present in C. chanardi, C. punctatonuchalis, C. siamensis, and C. vandeventeri). This species is distinguished from C. huaseesom by having keeled subtibial scales (vs smooth subtibial scales in C. huaseesom). This species is distinguished from C. siamensis and C. thachanaensis by lacking lineate gular marking (vs present in C. siamensis and C. thachanaensis). 
Comment 
EtymologyThe specific epithet refers to Adang and Rawi islands where the new species is found, and is a noun in apposition. 
References
  • Ampai N, Rujirawan A, Wood Jr PL, Stuart BL, Aowphol A 2019. Morphological and molecular analyses reveal two new insular species of Cnemaspis Strauch, 1887 (Squamata, Gekkonidae) from Satun Province, southern Thailand. ZooKeys 858: 127-161 - get paper here
  • Rujirawan, A., Yodthong, S., Ampai, N., Termprayoon, K., Aksornneam, A., Stuart, B. L., & Aowphol, A. 2022. A new rock gecko in the Cnemaspis siamensis group (Reptilia, Gekkonidae) from Kanchanaburi Province, western Thailand. Zoosystematics and Evolution, 98(2), 345-363 - get paper here
 
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