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Cnemaspis lineatubercularis AMPAI, WOOD, STUART & AOWPHOL, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Lan Saka Rock Gecko
Thai: Jing Jok Niew Yaow Lan Saka 
SynonymCnemaspis lineatubercularis AMPAI, WOOD, STUART & AOWPHOL 2020
Cnemaspis lineatubercularis — COTA et al. 2022 
DistributionThailand (Nakhon Si Thammarat)

Type locality: Thailand, Nakhon Si Thammarat Province, Lan Saka District, Kam Lon Subdistrict, Wang Mai Pak Waterfall (8°26.807'N, 99°46.525'E; 96 m a.s.l.)  
Reproduction 
TypesHolotype: ZMKU R 00828, adult male, collected on 25 January 2019 by Natee Ampai, Anchalee Aowphol, Attapol Rujirawan, Korkwan Termprayoon and Siriporn Yodthong.
Paratypes: Eighteen paratypes (adult males = 11, adult females = 7). ZMKU R 00821–00825 (five adult males), and ZMKU R 00826 (adult female), same data as holotype except that they were collected on 25 October 2016. ZMKU R 00827, ZMKU R 00829–00831 (four adult males), ZMKU R 00832–00835 (four adult fe- males), THNHM 28694–28695 (two adult males) and THNHM 28696–28697 (two adult females), same data as holotype. 
DiagnosisDiagnosis: Cnemaspis lineatubercularis sp. nov. can be distinguished from all other Cnemaspis by having the following combination of characters: (1) maximum SVL of 40.6 mm (mean 38.8 ± SD 1.4, N = 12) in adult males and maximum SVL of 41.8 mm (mean 39.5 ± SD 1.9, N = 7) in adult females; (2) 8–9 supralabial and infralabial scales; (3) gular, pectoral, abdominal, and subcaudal scales keeled; (4) rostral, interorbitals, supercilium, palmar scales, and ventral scales of brachia smooth; (5) 5–6 small, subconical spine-like tubercles present on flanks (6) 19–21 paravertebral tubercles linearly arranged; (7) 27–29 subdigital lamellae under the 4th toe; (8) 4–7 pore-bearing precloacal scales, pores rounded, arranged in chevron shape and separated in males; (9) one postcloacal tubercle each side in males; (10) ventrolateral caudal tubercles anteriorly present; (11) caudal tubercles restricted to a single paravertebral row on each side; (12) single median row of subcaudal scales keeled and lacking enlarged median row; and (13) gular region, abdomen, limbs and subcaudal region yellowish only in males. These differences are summarized among geographically close congeners in the siamensis group (Table 5).


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CommentHabitat: Specimens were found only along granitic rocky streams of Wang Mai Pak Waterfall. The rocky boulder microhabitats of this species are dry with cool surface temperatures (24.8–26.7 °C, 73.2–86.1% relative hu- midity). When disturbed, some individuals retreated deeper into rock crevices, cracks, more shaded areas or beneath rock boulders (Ampai et al.2020). 
EtymologyThe specific epithet lineatubercularis is taken from linea (Lat. for line) and tubercularis (Lat. for having tubercles), in reference to the new species having para-vertebral tubercles linearly arranged. 
References
  • Ampai N, Rujirawan A, Yodthong S, Termprayoon K, Stuart BL, Wood Jr PL, Aowphol A 2022. Hidden diversity of rock geckos within the Cnemaspis siamensis species group (Gekkonidae, Squamata): genetic and morphological data from southern Thailand reveal two new insular species and verify the phylogenetic affinities of C. chanardi and C. kam ZooKeys 1125: 115–158 - get paper here
  • Ampai N, Wood Jr PL, Stuart BL, Aowphol A 2020. Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159 - get paper here
  • Cota, M., Makchai, S., & Pongcharoen, C. 2022. Updated Catalogue of the Amphibian and Reptile Type Specimens of the Natural History Museum, National Science Museum, Thailand. Thai Specimens 21: 311––5407
  • Rujirawan, A., Yodthong, S., Ampai, N., Termprayoon, K., Aksornneam, A., Stuart, B. L., & Aowphol, A. 2022. A new rock gecko in the Cnemaspis siamensis group (Reptilia, Gekkonidae) from Kanchanaburi Province, western Thailand. Zoosystematics and Evolution, 98(2), 345-363 - get paper here
 
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