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Cnemaspis lineatubercularis AMPAI, WOOD, STUART & AOWPHOL, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Lan Saka Rock Gecko
Thai: Jing Jok Niew Yaow Lan Saka 
SynonymCnemaspis lineatubercularis AMPAI, WOOD, STUART & AOWPHOL 2020 
DistributionThailand (Nakhon Si Thammarat)

Type locality: Thailand, Nakhon Si Thammarat Province, Lan Saka District, Kam Lon Subdistrict, Wang Mai Pak Waterfall (8°26.807'N, 99°46.525'E; 96 m a.s.l.)  
Reproduction 
TypesHolotype: ZMKU R 00828, adult male, collected on 25 January 2019 by Natee Ampai, Anchalee Aowphol, Attapol Rujirawan, Korkwan Termprayoon and Siriporn Yodthong.
Paratypes: Eighteen paratypes (adult males = 11, adult females = 7). ZMKU R 00821–00825 (five adult males), and ZMKU R 00826 (adult female), same data as holotype except that they were collected on 25 October 2016. ZMKU R 00827, ZMKU R 00829–00831 (four adult males), ZMKU R 00832–00835 (four adult fe- males), THNHM 28694–28695 (two adult males) and THNHM 28696–28697 (two adult females), same data as holotype. 
DiagnosisDiagnosis. Cnemaspis lineatubercularis sp. nov. can be distinguished from all other Cnemaspis by having the following combination of characters: (1) maximum SVL of 40.6 mm (mean 38.8 ± SD 1.4, N = 12) in adult males and maximum SVL of 41.8 mm (mean 39.5 ± SD 1.9, N = 7) in adult females; (2) 8–9 supralabial and infralabial scales; (3) gular, pectoral, abdominal, and subcaudal scales keeled; (4) rostral, interorbitals, supercilium, palmar scales, and ventral scales of brachia smooth; (5) 5–6 small, subconical spine-like tubercles present on flanks (6) 19–21 paravertebral tubercles linearly arranged; (7) 27–29 subdigital lamellae under the 4th toe; (8) 4–7 pore-bearing precloacal scales, pores rounded, arranged in chevron shape and separated in males; (9) one postcloacal tubercle each side in males; (10) ventrolateral caudal tubercles anteriorly present; (11) caudal tubercles restricted to a single paravertebral row on each side; (12) single median row of subcaudal scales keeled and lacking enlarged median row; and (13) gular region, abdomen, limbs and subcaudal region yellowish only in males. These differences are summarized among geographically close congeners in the siamensis group (Table 5).

Comparisons. Cnemaspis lineatubercularis sp. nov. can be distinguished from other members of the siamensis group (C. adangrawi, C. chanardi, C. huaseesom, C. kamolnorranathi, C. omari, C. phangngaensis, C. punctatonuchalis, C. roticanai, C. siamensis, C. thachanaensis, and C. vandeventeri; Table 5) by having a smaller maximum SVL of 41.8 mm (vs. 44.9 mm in C. adangrawi, 43.5 mm in C. huaseesom, 42.0 mm in C. phangngaensis, 49.6 mm in C. punctatonuchalis, 47.0 mm in C. roticanai, 44.7 mm in C. vandeventeri) and by having a larger maximum SVL 41.8 mm (vs. 40.9 mm in C. chanardi, 37.8 mm in C. kamolnorranathi, 41.3 mm in C. omari, 39.7 mm in C. siamensis, 39.0 mm in C. thachanaensis).
Cnemaspis lineatubercularis sp. nov. is distinguished from C. adangrawi, C. phangngaensis, and C. thachanaensis by having fewer 8–9 supralabial scales (vs. 10 in C. adangrawi and C. phangngaensis, 10–11 in C. thachanaensis). This species is distinguished from C. phangngaensis by having fewer 8–9 infralabial scales (vs. 10 in C. phangngaensis). This species is distinguished from C. huaseesom and C. punctatonuchalis by having keeled ventral scales (vs. smooth ventral scales in C. huaseesom and C. punctatonuchalis). This species is distinguished from C. punctatonuchalis, C. siamensis, and C. thachanaensis by presence of precloacal pores (vs. precloacal pores absent in C. punctatonuchalis, C. siamensis, C. thachanaensis). This species is distinguished from C. huaseesom, C. kamolnorranathi, and C. phangngaensis by having a separated row of precloacal pores (vs. continuous in C. huaseesom, C. kamolnorranathi, C. phangngaensis). This species is distinguished from C. kamolnorranathi by having rounded precloacal pores (vs. pores elongated in C. kamolnorranathi).
Cnemaspis lineatubercularis sp. nov. is distinguished from C. adangrawi, C. chanardi, C. omari, C. phangngaensis, C. punctatonuchalis, C. roticanai, and C. vandeventeri by having fewer 19–21 paravertebral tubercles (vs. 23–25 in C. adangrawi, 22–25 in C. chanardi, 22–29 in C. omari, 22 in C. phangngaensis, 24–27 in C. punctatonuchalis, 25–27 in C. roticanai, 25–29 in C. vandeventeri). This species is distinguished from C. adangrawi, C. chanardi, C. huaseesom, C. omari, C. roticanai, C. siamensis, and C. vandeventeri by having paravertebral tubercles linearly arranged (vs. randomly arranged in C. adangrawi, C. chanardi, C. huaseesom, C. omari, C. roticanai, C. siamensis, C. vandeventeri). This species is distinguished from C. adangrawi and C. phangngaensis by having tubercles on lower flanks (vs. absent in C. adangrawi and C. phangngaensis). This species is distinguished from C. siamensis and C. thachanaensis by having more 27–29 lamellae under 4th toe (vs. 24–26 in C. siamensis and 24 in C. thachanaensis).
Cnemaspis lineatubercularis sp. nov. is distinguished from C. chanardi, C. huaseesom, C. kamolnorranathi, C. omari, C. roticanai, C. siamensis, and C. vandeventeri by the presence of ventrolateral caudal tubercles anteriorly (vs. lacking in C. chanardi, C. huaseesom, C. kamolnorranathi, C. omari, C. roticanai, C. siamensis, C. vandeventeri). This species is distinguished from C. vandeventeri by having lateral caudal furrows (vs. lacking in C. vandeventeri). This species is distinguished from C. huaseesom and C. punctatonuchalis by having keeled subcaudal scales (vs. lacking in C. huaseesom and C. punctatonuchalis). This species is distinguished from C. chanardi, C. huaseesom, C. omari, C. punctatonuchalis, C. roticanai, and C. siamensis by having single median row of keeled subcaudals (vs. lacking in C. chanardi, C. huaseesom, C. omari, C. punctatonuchalis, C. roticanai, C. siamensis). This species is distinguished from C. chanardi, C. punctatonuchalis, C. siamensis, and C. vandeventeri by lacking enlarged median subcaudal scales (vs. present in C. chanardi, C. punctatonuchalis, C. siamensis, C. vandeventeri). This species is distinguished from C. adangrawi, C. chanardi, C. huaseesom, C. kamolnorranathi, C. omari, C. punctatonuchalis, C. roticanai, C. siamensis, and C. vandeventeri by having caudal tubercles restricted to a single paravertebral row on each side (vs. lacking in C. adangrawi, C. chanardi, C. huaseesom, C. kamolnorranathi, C. omari, C. punctatonuchalis, C. roticanai, C. siamensis, C. vandeventeri).
Cnemaspis lineatubercularis sp. nov. is distinguished from C. thachanaensis by having one postcloacal tubercle in males (vs. lacking in C. thachanaensis). This species is distinguished from C. huaseesom by having keeled subtibial scales (vs. smooth in C. huaseesom). This species is distinguished from C. adangrawi, C. kamolnorranathi, C. punctatonuchalis, C. siamensis, C. thachanaensis, and C. vandeventeri by having yellow coloration in the subcaudal region (vs. lacking in C. adangrawi, C. kamolnorranathi, C. punctatonuchalis, C. siamensis, C. thachanaensis, C. vandeventeri). This species is distinguished from C. huaseesom, C. phangngaensis, C. punctatonuchalis, and C. roticanai by lacking dorsal color pattern sexually dimorphic (vs. having in C. huaseesom, C. phangngaensis, C. punctatonuchalis, C. roticanai). 
CommentHabitat: Specimens were found only along granitic rocky streams of Wang Mai Pak Waterfall. The rocky boulder microhabitats of this species are dry with cool surface temperatures (24.8–26.7 °C, 73.2–86.1% relative hu- midity). When disturbed, some individuals retreated deeper into rock crevices, cracks, more shaded areas or beneath rock boulders (Ampai et al.2020). 
EtymologyThe specific epithet lineatubercularis is taken from linea (Lat. for line) and tubercularis (Lat. for having tubercles), in reference to the new species having para- vertebral tubercles linearly arranged. 
References
  • Ampai N, Wood Jr PL, Stuart BL, Aowphol A 2020. Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159 - get paper here
 
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