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Coelognathus subradiatus (SCHLEGEL, 1837)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Sunda Rat Snake
G: Indonesische Kletternatter 
SynonymColuber subradiatus SCHLEGEL 1837: 136
Compsosoma sub-radiatum — DUMÉRIL, BIBRON & DUMÉRIL 1854: 297
Coluber subradiatus — BOULENGER 1894: 64
Elaphis subradiatus — LIDTH DE JEUDE 1895: 126
Coluber subradiatus — BOULENGER 1897: 506
Coluber melanurus var. timoriensis FERREIRA 1897 (fide KAISER, pers. comm.)
Elaphe subradiata — SCHULZ 1996: 247
Coelognathus subradiatus — HELFENBERGER 2001
Coelognathus subradiata — PYRON & BURBRINK 2013
Coelognathus subradiatus — WALLACH et al. 2014: 172 
DistributionIndonesia (Enggano, Lombok, Sumbawa, Sumba, Komodo, Flores, Alor, Roti, Semau, Timor, Wetar, Timor-Leste

Type locality: Indonesia: Timor  
Reproductionoviparous 
TypesType: BMNH 1946.1.6.16 (and possibly additional specimens). 
Diagnosis 
CommentSynonymy: "Based on the original description, Coluber melanurus var. timoriensis BETHENCOURT FERREIRA 1897 can be assigned to Coelognathus subradiatus and not C. flavolineatus. Unfortunately the specimens used by Bethencourt Ferreira were destroyed in the 1978 fire at the Museu Bocage in Lisbon, so this assessment relies on the comparison of new material from Timor-Leste with Bethencourt Ferreira's description” (H. Kaiser, pers. comm.)

Subspecies: Coelognathus subradiatus enganensis (VINCIGUERRA 1892) has been elevated to full species status. 
References
  • Boulenger, G. A. 1898. A list of reptiles and batrachians from Ombaai, East Indian Archipelago. Ann. Mag. Nat. Hist. (7) 1: 122-124 - get paper here
  • Boulenger, G.A. 1897. List of the reptiles and batrachians collected by Mr. Alfred Everett in Lombok, Flores, Sumba and Saru, with descriptions of new species. Ann. Mag. Nat. Hist. (6) 19: 503-509 - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Ferreira, J. Bethencourt 1897. Sobre alguns reptis ultimamente enviados á secção zoologica do Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Lisboa 5 (18): 111-116
  • Helfenberger, Notker 2001. Phylogenetic relationship of Old World Ratsnakes based on visceral organ topography, osteology, and allozyme variation. Russ. J. Herpetol. (Suppl.): 1-56 - get paper here
  • KENNEDI, UMAR F.; MIRZA D. KUSRINI, ANI MARDIASTUTI, AND ACHMAD ARIEFIANDY 2021. THE AMPHIBIANS AND REPTILES IN KOMODO NATIONAL PARK AND THE SURROUNDING AREA. Media Konservasi 26 (3): 217-230; DOI: 10.29244/medkon.26.3.217-230 - get paper here
  • Knight, Sue 2009. Some Captive Observation Notes on The Timor Ratsnake - Coelognathus subradiatus subradiatus. Litteratura Serpentium 29 (4): 156-165 - get paper here
  • Lang, Ruud de 2011. The Snakes of the Lesser Sunda Islands (Nusa Tenggara), Indonesia. Asian Herpetological Research 2 (1): 46-54 - get paper here
  • Lang, Ruud de 2012. Snakes of the Lesser Sunda Islands (Nusa Tenggara), Indonesia. Edition Chimaira, 349 pp. - get paper here
  • Lidth de Jeude, Th. W. van. 1895. Reptiles from Timor and the neighbouring islands. Notes from the Leyden Museum 16: 119-127 - get paper here
  • Mecke, Sven 2016. Unbekannte Artenvielfalt in einer „vergessenen Welt“: die Herpetofauna von Ost-Timor. Terraria-Elaphe 2016 (4): 52-55 - get paper here
  • Mertens, R. 1930. Die Amphibien und Reptilien der Inseln Bali, Lombok, Sumbawa und Flores. Senck. Naturf. Gesell., Frankfurt am Main, Abhandl. 42(3): 117-344.
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Andrew KATHRINER, Sven MECKE, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2015. Herpetological Diversity of Timor-Leste: Updates and a Review of Species Distributions. Asian Herpetological Research 6 (2): 73–131 - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Scott HEACOX, Andrew KATHRINER, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2012. First Update to Herpetofaunal Records from Timor-Leste. Asian Herpetological Research 3 (2): 114–126 - get paper here
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • Reilly, Sean B.; Alexander L. Stubbs, Benjamin R. Karin, Umilaela Arifin, Evy Arida, Djoko T. Iskandar, and Jimmy A. McGuire 2021. Genetic divergence of the Sunda ratsnake (Coelognathus subradiatus) across the Lesser Sunda Islands (Squamata: Colubridae). Amphibia-Reptilia 42 (2): 269–273 - get paper here
  • Schlegel, H. 1837. Essai sur la physionomie des serpens. Partie Descriptive. La Haye (J. Kips, J. HZ. et W. P. van Stockum), 606 S. + xvi - get paper here
  • Schlegel, H. 1837. Abbildungen neuer oder unvollständig bekannter Amphibien, nach der Natur oder dem Leben entworfen. Düsseldorf (Arnz & Comp.), i-xiv + 141 pp. [1837-1844] - get paper here
  • Schulz, K. D. 1996. Eine Monographie der Schlangengattung Elaphe Fitzinger. Bushmaster, Berg (CH): 1-460
  • Schulz, Klaus Dieter 1988. Die hinterasiatischen Kletternattern der Gattung Elaphe. Teil XIV Elaphe subradiata (Schlegel 1837) und Elaphe erythrura (Duméril & Bibron 1854). Sauria 10 (3): 17-20 - get paper here
  • Schulz, Klaus-Dieter 1996. A monograph of the colubrid snakes of the genus Elaphe Fitzinger. Koeltz Scientific Books, 439 pp.
  • Suyanto, R.L.A.A.H. 1996. Geographical Variation in the Morphology of Four Snake Species from the Lesser Sunda Islands, Eastern Indonesia. Biological Journal of the Linnean Society 59(4):439-456. - get paper here
  • Vinciguerra, D. 1892. Rettili e batraci di Engano. Ann. Mus. civ. stor. nat. Genova, 2. Ser. 7 (32): 517-526. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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