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Complicitus nigrigularis (OTA & HIKIDA, 1991)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Blackthroated Bloodsucker 
SynonymCalotes nigrigularis OTA & HIKIDA 1991
Bronchocela nigrigularis – MALKMUS 1994: 235
Complicitus nigrigularis — MANTHEY & GROSSMANN 1997: 166
Complicitus nigrigularis — BARTS & WILMS 2003
Complicitus nigrigularis — DAS & YAAKOB 2007 
DistributionMalaysia (Borneo, Sabah)

Type locality: Kinabalu Park, Sabah, Borneo.  
Reproductionoviparous 
TypesHolotype: KUZ 13223, adult male 
DiagnosisDiagnosis (genus): A relatively small agamid lizard with a snout-vent length (SVL) of < 80 mm and a tail about twice as long, which resembles Pseudocalotes species in general habitus. Easily distinguishable from these and all other agamid lizard genera by the combination of the following characters: gular sac with lateral pockets and granulated scales, crests on neck, back and along the median line of the gular pouch present, dorsals heterogeneous and tympanum visible (Manthey & Denzer 2019: 8). 
CommentType species: Calotes nigrigularis OTA & HIKIDA 1991 is the type species of the genus Complicitus MANTHEY & GROSSMANN 1997.

Abundance: only known from the type locality (Meiri et al. 2017) and from 3 specimens (Manthey & Denzer 2019). 
EtymologyNamed after the black coloration of the throat, from Latin niger, nigra, nigrum = black, and Latin gula = throat, also for esophagus. 
References
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. Lakim, M. 2008. A second specimen of Complicitus nigrigularis (Ota & Hikida, 1991). Hamadryad 32 (1): 71-73 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Malkmus,R. 1994. Herpetologische Beobachtungen aam Mt. Kinabalu, Nord-Borneo. IV. Mitt. Zool. Mus. Berlin 70: 217-249. - get paper here
  • MANTHEY, U. & DENZER, W. 2019. Rare mountain agamas of the Kinabalu Park (Sabah, North Borneo) above 900 m a.s.l. Sauria 41 (1): 2 - 12 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Ota H; Hikida T 1991. Taxonomic review of the lizards of the genus Calotes Cuvier 1817 (Agamidae Squamata) from Sabah, Malaysia. Tropical Zoology 4 (2): 179-192 - get paper here
 
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