Cordylus machadoi LAURENT, 1964

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Higher Taxa	Cordylidae (Cordylinae), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Machado’s Girdled Lizard, Kaokoveld Girdled Lizard G: Machado’s Gürtelschweif
Synonym	Cordylus vittifer machadoi LAURENT 1964: 49 Cordylus machadoi — BRANCH 1998 Cordylus machadoi — STANLEY et al. 2011 Cordylus machadoi — REISSIG 2014 Cordylus machadoi — SCHLEICHER 2020
Distribution	S Angola, N Namibia Type locality: Leba, Humpata, environs de Sá da Bandeira, elevation 1800 m, Huila.
Reproduction	Ovoviviparous.
Types	Holotype: MD = Museu Dundo; paratype: MCZ
Diagnosis	Diagnosis: “Race de Cordylus vittifer différant de la forme typique par son plus grand nombre de rangées longitudinales d'écailles dorsales: 26-28 au lieu de 20-24 et la rostrale moins large (hauteur comprise 2,34 à 2,32 fois dans sa largeur au lieu de 3 à 4 fois)” (Laurent 1964).
Comment	Distribution: see map in GREENBAUM et al. 2012. Cordylus v. machadoi was originally distinguished from other C. vittifer on the basis of its greater numbers of longitudinal dorsal scale rows (26–28 versus 20–24) and a narrower rostral scale (2.32–2.34 times wider than deep versus 3–4 times wider). Reichenow (1887), in his original description of Zonurus vittifer (Fig. S2), reported only 20–22 longitudinal dorsal scale rows and did not mention the size or shape of the rostral, and it therefore seems that Laurent (1964) was referring to Loveridge (1944a, see table on p. 15 in Bates et al. 2023) and possibly also FitzSimons (1943) when comparing machadoi to vittifer.
Etymology	Named after Dr. A. de Barros-Machado, who collected the type in Angola.
References	Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here Bates MF, Lobón-Rovira J, Stanley EL, Branch WR, Vaz Pinto P 2023. A new species of green-eyed Cordylus Laurenti, 1768 from the west-central highlands of Angola, and the rediscovery of Cordylus angolensis (Bocage, 1895) (Squamata: Cordylidae). Vertebrate Zoology 73: 599-646 - get paper here Branch, W. R. 1998. Field Guide to the Snakes and Other Reptiles of Southern Africa. 3rd ed. Fully Revised and Updated to Include 83 New Species. Ralph Curtis Books (Sanibel Island, Florida), 399 pp. Butler, B O; Ceríaco, L M P; Marques, M P; Bandeira, S; Júlio, T; Heinicke, M P & Bauer, A M; 2019. Herpetological survey of Huíla Province, Southwest Angola, including first records of Bicuar National Park. Herpetological Review 50 (2): 225-240 - get paper here Ceríaco, Luis M. P.; Mariana P. Marques, Ilunga André, Esteves Afonso, David C. Blackburn & Aaron M. Bauer 2020. ILLUSTRATED TYPE CATALOGUE OF THE “LOST” HERPETOLOGICAL COLLECTIONS OF MUSEU DO DUNDO, ANGOLA. Bulletin of the Museum of Comparative Zoology 162 (7): 379-440 - get paper here Conradie, Werner; Brain Reeves, Sandile Mdoko, Lwandiso Pamla, Oyama Gxabhu 2020. Herpetological survey of the Ongeluksnek (Malekgalonyane) Nature Reserve on the foothills of the Drakensberg, Eastern Cape Province, South Africa. Herpetology Notes 13: 717-730 - get paper here Greenbaum, Eli; Edward L. Stanley, Chifundera Kusamba, Wandege M. Moninga, Stephen R. Goldberg & Charles R. Bursey 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61 (1): 14-39 - get paper here Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here Laurent, R.F. 1964. Reptiles et batraciens de l'Angola (troisième contribution). Companhia de Di amantes de Angola (Diamang), Serviços Culturais, Museu do Dundo (Angola), No. 67, 165 pp. Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II) MARQUES, MARIANA P.; LUIS M. P. CERÍACO, EDWARD L. STANLEY, SUZANA A. BANDEIRA, ISHAN AGARWAL, AARON M. BAUER 2019. A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola. Zootaxa 4668 (4): 503–524 - get paper here Reissig, J. 2014. Girdled Lizards and Their Relatives. Edition Chimaira, Frankfurt, ISBN-10: 3899734378 Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp. Stanley, Edward L.; Aaron M. Bauer; Todd R. Jackman, William R. Branch, P. Le Fras N. Mouton 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): 53-70. - get paper here STANLEY, EDWARD L.; LUIS M. P. CERÍACO, SUZANA BANDEIRA, HILARIA VALERIO, MICHAEL F. BATES & WILLIAM R. BRANCH 2016. A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert,. Zootaxa 4061 (3): 201–226 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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