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Crotalus pusillus KLAUBER, 1952

IUCN Red List - Crotalus pusillus - Endangered, EN
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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Tancitaran Dusky Rattlesnake
S: Cascabel Oscura del Tancítaro 
SynonymCrotalus pusillus KLAUBER 1952: 34
Crotalus pusillus — LINER 1994
Crotalus pusillus — MCDIARMID, CAMPBELL & TOURÉ 1999: 292
Uropsophus pusillus — HOSER 2009
Crotalus pusillus — WALLACH et al. 2014: 193 
DistributionWC Mexico (NE Michoacan, S Jalisco)

Type locality: Michoacan, Tancitaro; 5,000 feet elevation  
Reproductionovovivparous 
TypesHolotype: FMNH 39112; paratype: MSNG 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2471 characters), are available for collaborators and contributors. Please contact us for details. 
CommentVenomous!

Nomenclature: Hoser’s 2009 classification and nomenclature has been rejected as unnecessary and unavailable by WÜSTER & BERNILS 2011.

Conservation: one of the 30 most endangered viper species (Maritz et al. 2016).

Distribution: see map in Blair et al. 2018: 352 (Fig. 1). 
EtymologyNamed after Latin “pusillus, -a, -um” = small, tiny. 
References
  • Blair C, Bryson RW Jr, Linkem CW, Lazcano D, Klicka J, McCormack JE. 2018. Cryptic diversity in the Mexican highlands: Thousands of UCE loci help illuminate phylogenetic relationships, species limits and divergence times of montane rattlesnakes (Viperidae: Crotalus). Molecular Ecology Resources 19: 349–365 [2019] - get paper here
  • Bryson Jr, Robert W.; Robert W. Murphy, Amy Lathrop and David Lazcano-Villareal 2011. Evolutionary drivers of phylogeographical diversity in the highlands of Mexico: a case study of the Crotalus triseriatus species group of montane rattlesnakes. Journal of Biogeography 38: 697–710 - get paper here
  • Bryson, R.W. 2007. Fotopirsch auf Gebirgsklapperschlangen. Reptilia (Münster) 12 (66): 32-37 - get paper here
  • BRYSON, ROBERT W.; JR., CHARLES W. LINKEM, MICHAEL E. DORCAS, AMY LATHROP, JASON M. JONES, JAVIER ALVARADO-DÍAZ, CHRISTOPH I. GRÜNWALD & ROBERT W. MURPHY 2014. Multilocus species delimitation in the Crotalus triseriatus species group (Serpentes: Viperidae: Crotalinae), with the description of two new species. Zootaxa 3826 (3): 475–496 - get paper here
  • Campbell, J.A. & Lamar, W.W. 1989. The Venomous Reptiles of Latin America. Comstock Publishing/Cornell University Press, Ithaca
  • Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp
  • Hoser, R. 2009. A reclassification of the rattlesnakes; species formerly exclusively referred to the genera Crotalus and Sistrurus. Australasian J. Herpetol. 3: 1-21 - get paper here
  • Klauber, Laurence M. 1952. Taxonomic studies on rattlesnakes of Mainland Mexico. Bulletins of the Zoological Society of San Diego (26): 1-143 - get paper here
  • Lemos-Espinal JA, Smith GR, Pierce LJS, Painter CW 2020. The amphibians and reptiles of Colima, Mexico, with a summary of their conservation status. ZooKeys 927: 99-125 - get paper here
  • Maritz, Bryan; Johannes Penner, Marcio Martins, Jelka Crnobrnja-Isailović, Stephen Spear, Laura R.V. Alencar, Jesús Sigala-Rodriguez, Kevin Messenger, Rulon W. Clark, Pritpal Soorae, Luca Luiselli, Chris Jenkins, Harry W. Greene 2016. Identifying global priorities for the conservation of vipers. Biological Conservation, doi:10.1016/j.biocon.2016.05.004 - get paper here
  • McCranie J R 1983. Crotalus pusillus Klauber. Southwestern Mexico dusky rattlesnake. Catalogue of American Amphibians and Reptiles ( 313: 1-2 - get paper here
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp.
  • Meik, Jesse M and André Pires-daSilva 2009. Evolutionary morphology of the rattlesnake style. BMC Evolutionary Biology 2009, 9:35 - get paper here
  • Muñoz-Nolasco, Francisco Javier; Daniel Cruz-Sáenz, Oscar Jair Rodríguez-Ruvalcaba and Isaac Enrique Terrones-Ferreiro 2015. Notes on the Herpetofauna of Western Mexico 12: Herpetofauna of a Temperate Forest in Mazamitla, Southeastern Jalisco, Mexico. Bull. Chicago Herp. Soc. 50 (4):45-50 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wüster, W. & Bérnils, R.S. 2011. On the generic classification of the rattlesnakes, with special reference to the Neotropical Crotalus durissus complex (Squamata: Viperidae). ZOOLOGIA 28 (4): 417–419 - get paper here
 
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