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Cubatyphlops notorachius (THOMAS & HEDGES, 2007)

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Higher TaxaTyphlopidae (Typhlopinae), Typhlopoidea, Serpentes, Squamata (snakes)
Common NamesE: Imias Blindsnake 
SynonymTyphlops notorachius THOMAS & HEDGES 2007: 21
Cubatyphlops notorachius — HEDGES et al. 2014
Typhlops notorachius — PYRON & WALLACH 2014
Cubatyphlops notorachius — NAGY et al. 2015
Typhlops notorachius — WALLACH et al. 2014: 768 
DistributionE Cuba (east of Guantánamo Bay)

Type locality: 9.4 km W Imias, Guantánamo Province, Cuba, 5 m elevation.  
TypesHolotype: MNHNCu 4551 (field tag number 191322), a male, collected on 1 July 1990 by S. Blair Hedges, Richard Thomas, and Daniel McCallister. 
DiagnosisDiagnosis: A relatively large species of the Typhlops biminiensis group differing from T. biminiensis in having a rostral with an acuminate posterior edge, not broadly rounded. Also, the rostral has parallel sides and is not indented at apex of the snout (Fig. 8). Additionally, T. notorachius has a proportionately smaller rostral than the preceding species (T. biminiensis, T. arator, T. perimychus, and T. anousius) (Figs. 8, 10A). The difference in body size between T. notorachius and T. perimychus is striking; not considering the associated specimen of the latter species (which may be a different species), all four specimens of T. notorachius (282– 301 mm TL) are larger than all 18 specimens of T. perimychus (130–280 mm TL). Because each species was collected at multiple localities and, in the case of T. perimychus, over four decades, we surmise that the body size difference is real and not the result of collecting bias. Also, T. notorachius differs from T. perimychus in having a greater anterior nasal width (ANTNAS/RW1): 0.42–0.53 versus 0.34–0.39 in T. perimychus. From T. anousius, T. notorachius also differs in its point of scale row reduction: 15–40% TL versus 2% TL in T. anousius. 
EtymologyAn adjective made from the Greek, notos, south, and rhachia, shore, meaning “of the southern shore.” 
  • DOMÍNGUEZ, MICHEL & LUIS V. MORENO 2009. Taxonomy of the Cuban blind snakes (Scolecophidia, Typhlopidae), with the description of a new large species. Zootaxa 2028: 59-66 - get paper here
  • Hedges, S.B., Marion, A.B., Lipp, K.M., Marin, J. & Vidal, N. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49: 1–61 - get paper here
  • Kornilios, P.; S. Giokas, P. Lymberakis, R. Sindaco 2013. Phylogenetic position, origin and biogeography of Palearctic and Socotran blind-snakes (Serpentes: Typhlopidae). Molecular Phylogenetics and Evolution 68 (1): 35–41 - get paper here
  • NAGY, ZOLTÁN T.; ANGELA B. MARION, FRANK GLAW, AURÉLIEN MIRALLES,<br />JOACHIM NOPPER, MIGUEL VENCES & S. BLAIR HEDGES 2015. Molecular systematics and undescribed diversity of Madagascan scolecophidian snakes (Squamata: Serpentes). Zootaxa 4040 (1): 031–047 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Pyron, R.A. & Wallach, V. 2014. Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa 3829 (1): 001–081 - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • THOMAS R. & S.B. HEDGES 2007. Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba. Zootaxa 1400: 1-26 - get paper here
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