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Cyrtodactylus bokorensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus bokorensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER 2019
Gymnodactylus intermedius SMITH 1935: 44
Cyrtodactylus intermedius STUART & EMMETT 2006: 17 
DistributionCambodia (Kampot)

Type locality: Bokor National Park Field Station, Kampot Province, Cambodia (10 37.611'N, 104 01.541'E); 1033 m elevation  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 8554, adult male, collected on 10 July 2007 by L. Lee Grismer and Neang Thy. Paratypes. Adult males LSUHC 8541–42, adult females LSUHC 8548–50 and 8555 bear the same collection data as the holotype. 
DiagnosisDiagnosis: Adult males reaching 81.9 mm SVL, adult females reaching 93 mm SVL; eight supralabials, 8–10 infralabials; 30–33 paravertebral tubercles; 18–20 longitudinal rows of dorsal tubercles; 42–48 rows of ventral scales; six or seven expanded subdigital lamellae proximal to the digital inflection, 11–14 unmodified, distal, subdigital lamellae; 18–20 total subdigital lamellae on fourth toe; enlarged femoral and precloacal scales vary from being separated by a diastema to continuous; 26–30 enlarged femoral scales; proximal femoral scales less than half the size of distal proximal scales; 7–10 enlarged precloacal scales with pores on each in males; two or three rows of enlarged post-precloacal scales; two or three postcloacal tubercles; no pocketing present between digits of hind or forefeet; dark pigmented blotches on top of head vary from present to absent; posterior border of nuchal loop pointed, chevron-shaped; and four or five dark body bands (summarized in Tables 7 + 10 in Murdoch et al. 2019).


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CommentHabitat: scattered sandstone rocks and boulders among highly disturbed wind-blown scrub vegetation. Also abundant at 350 m elevation in wet evergreen forest associated with rocky areas on the southern side of Bokor NP and therefore probably common throughout Bokor National Park. 
EtymologyThe specific epithet, bokorensis sp. nov., is an adjective in reference to type locality of the Bokor Plateau to which it is presumably endemic. 
References
  • CHHIN, S., NEANG, T., CHAN, S., KONG, K., OU, R., SAMORN, V., ... & GRISMER, L. L. 2024. A new species in the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from an isolated limestone karst formation in southwestern Cambodia. Zootaxa 5474 (1): 1-20
  • GRISMER, L. LEE; CHAN KIN ONN, JAMIE R. OAKS, THY NEANG, LANG SOKUN,<br>MATTHEW L. MURDOCH, BRYAN L. STUART & JESSE L. GRISMER 2020. A new insular species of the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from Cambodia with a discussion of habitat preference and ecomorphology. Zootaxa 4830 (1): 075–102 - get paper here
  • MURDOCH, MATTHEW L.; L. LEE GRISMER, PERRY L. JR. WOOD, THY NEANG, NIKOLAY A. POYARKOV, NGO VAN TRI, ROMAN A. NAZAROV, ANCHALEE AOWPHOL, OLIVIER S.G. PAUWELS, HUNG NGOC NGUYEN, JESSE L. GRISMER 2019. Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa 4554 (1): 001–062 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Stuart, B.L. & Emmett, D.A. 2006. A Collection of Amphibians and Reptiles from the Cardamom Mountains, Southwestern Cambodia. Fieldiana Zool. N.S. (109): 1-27 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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