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Cyrtodactylus calamei LUU, BONKOWSKI, NGUYEN, LE, SCHNEIDER, NGO & ZIEGLER, 2016

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Calame’s Bent-toed Gecko
Laotian: Ki Chiem Calame 
SynonymCyrtodactylus calamei LUU, BONKOWSKI, NGUYEN, LE, SCHNEIDER, NGO & ZIEGLER 2016 
DistributionC Laos

Type locality: karst forest, Tham Nok Aen region, Thong Xam Village (17°34.179’N, 105°50.329’E, elevation 210 m a.s.l.) within Hin Nam No NPA, Khammouane Province, central Laos.  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: VNUF R.2015.28, adult male, collected on 25 March 2015 by V. Q. Luu, T. Calame, and K. Thanabuaosy. Paratypes. IEBR A. 2015.36, adult male; NUOL R-2015.22, subadult male; VNUF R.2015.27, adult female, the same data as the holotype. 
DiagnosisDiagnosis: Cyrtodactylus calamei sp. nov. can be distinguished from its congeners by a unique combination of the following characters: Adult SVL 80.0 ± 8.0 mm (mean ± SD); head dorsally with grey small scattered spots; nuchal loop present with indentations, not enlarged posteriorly, extending from each postnasal cross orbit and contacting on nape; four greyish brown, wide transverse bands between limbs, sometimes irregular; dorsal surface with homogenous, low, round, weakly keeled scales; 39–42 ventral scales at midbody; ventrolateral skin folds well-defined; 183–197 ventral scale rows from mental to cloacal slit; 101–114 scale rows at midbody; 35–39 precloacal-femoral pores in males, 38 in the female; enlarged femoral and precloacal scales present; four postcloacal tubercles; subcaudal scales transversely enlarged.


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CommentSexual dimorphism. The single adult female differs from two adult males by its larger size (maximum SVL 89.3 mm versus 75.8 mm in the males) and lacking of hemipenial swellings at the base of tail (see Table 4 & Fig. 6, Luu et al. 2016).

Natural history. Specimens were found at night between 19:30 and 21:08h, on limestone outcrops, at elevations between 190 and 260 m a.s.l. The surrounding habitat was karst forest. The relative humidity was 80% and the air temperature ranged from 23 to 26oC (see Table 5, Luu et al. 2016).

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyNamed after Mr. Thomas Calame, from WWF Greater Mekong, Vientiane, Laos, who participated in the field research of the authors in Hin Nam No NPA, Khammouane Province between 2014 and 2015. 
References
  • Kwet, Axel 2017. Neue Arten: Liste der im Jahr 2016 neu beschriebenen Reptilien. Terraria-Elaphe 2017 (3): 54-70 - get paper here
  • LUU, VINH QUANG; MICHAEL BONKOWSKI, Truong QUang Nguyen, MINH DUC LE, NICOLE SCHNEIDER, HANH THI NGO, THOMAS ZIEGLER 2016. Evolution in karst massifs: Cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos. Zootaxa 4107 (2): 101–140 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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