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Cyrtodactylus naungkayaingensis GRISMER, WOOD, THURA, QUAH, MURDOCH, GRISMER, HERR, LIN & KYAW, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus naungkayaingensis GRISMER, WOOD, THURA, QUAH, MURDOCH, GRISMER, HERR, LIN & KYAW 2018 
DistributionMyanmar (Kayin: Hpa-an District)

Type locality: Naung Ka Yaing Hill, 17 km north-east of Hpa-an, Hpa-an District, Kayin State, Myanmar (17.01558°N, 97.70641°E; 14 m in elevation.  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 13209, Adult male, collected on 8 May 2017 at 2000 hrs by Myint Kyaw Thura, Mark W. Herr, L. Lee Grismer, Perry L. Wood, Jr., Matthew L. Murdoch, Evan S.H. Quah, and Htet Kyaw. Paratypes: Adult male LSUHC 13208 and juvenile LSUHC 13207 bear the same collecting data as the holotype. 
DiagnosisDiagnosis: Cyrtodactylus naungkayaingensis sp. nov. differs from all congeners by having the unique combination of maximum SVL 66.9 mm; eight or nine supralabials; seven infralabials; 34 or 35 paravertebral tubercles; 16–18 longitudinal rows of body tubercles; 27 longitudinal rows of ventral scales; relatively long digits with seven expanded subdigital lamellae proximal to the digital inflexion on the fourth toe, 11 or 12 unmodified distal subdigital lamellae, 18–19 total subdigital lamellae; raised, moderately keeled, dorsal body tuber- cles in adults; tubercles extending beyond base of tail; enlarged femoral and precloacal scales continuous; enlarged proximal femoral scales less than one-half the size of enlarged distal femoral scales; 30–35 enlarged femoral scales; 11 femoral pores in males; three or four precloacal pores in males; pore-bearing femoral and precloacal scales discontinuous; 8–12 enlarged precloacal scales; three rows of enlarged post- precloacal scales; medial subcaudal scales 3 times as wide as long, extending onto lateral surface of tail; top of head mottled in adults, no yellow reticulum; nuchal loop not divided medially, no pronounced anterior azygous notch, posterior border jagged; four jagged, dark, dorsal bands with lightened centres bearing paravertebral elements, narrower than interspaces, not edged with white tubercles; nape band variably present; dark markings in dorsal interspaces; ventrolateral folds not whitish; anterodorsal margins of thighs and brachia not pigmented; 11 light-coloured caudal bands bearing dark markings, not encircling tail; 10 dark caudal bands wider than light caudal bands; and mature regenerated tail not spotted. These characters are scored against all other species in the sinyineensis group in Table 4 and all other species of the Indochinese clade in Grismer et al. (2017a, table 8).


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Comment 
EtymologyThe specific epithet, naungkayaingensis, is a noun in apposition in reference to the type locality of Naung Ka Yaing Hill. 
References
  • CHOMDEJ, S., SUWANNAPOOM, C., PAWANGKHANANT, P., PRADIT, W., NAZAROV, R. A., GRISMER, L. L., & POYARKOV, N. A. 2020. A new species Cyrtodactytlus Gray (Squamata: Gekkonidae) from western Thailand and the phylogenetic placement of C. inthanon and C. doisuthep. Zootaxa 4838 (2): 179-209 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr., Myint Kyaw Thura, Evan S.H. Quah, Matthew L. Murdoch, Marta S. Grismer, Mark W. Herr, Aung Lin & Htet Kyaw 2018. Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History, 52:19-20, 1243-1294 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S.H. QUAH, MARTA S. GRISMER, MYINT KYAW<br>THURA, JAMIE R. OAKS & AUNG LIN 2020. Two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from a karstic archipelago in the Salween Basin of southern Myanmar (Burma). Zootaxa 4718 (2): 151–183 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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