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Cyrtodactylus soni LE, NGUYEN, LE & ZIEGLER, 2016

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Son’s Bent-toed Gecko (English)
Vietnamese: Thạch sùng ngón sơn 
SynonymCyrtodactylus soni LE, NGUYEN, LE & ZIEGLER 2016 
DistributionVietnam (Ninh Binh)

Type locality: karst forest near Da Han Village (20o25.067’N, 105o51.467’E, elevation 17 m a.s.l.), Gia Hoa Commune, within Van Long Wetland Nature Reserve, Gia Vien District, Ninh Binh Province, Vietnam  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: HNUE VL.2015.78, adult male, collected on 24 July 2015 by D. T. Le, A. M. Luong, D. T. Pham, and N. H. Nguyen. Paratypes. IEBR R.2016.4, adult female, HNUE VL.2015.94, adult female, collected on 24 July 2015; IEBR R.2016.5, adult male, HNUE VL.2015.131, adult female, and HNUE VL.2015.132, adult female, collected on 18 August 2015, the same data as the holotype. 
DiagnosisDiagnosis: The new species can be distinguished from other members of the genus Cyrtodactylus from Indochina by a combination of the following characters: medium size (SVL up to 103 mm); internasal single; dorsal tubercles in 10–13 irregular rows; ventral scale rows 41–45; lateral skin folds present, without interspersed tubercles; precloacal pores 6 or 7 in males, 7 or 8 pitted scales in females, in a continuous row; femoral pores 6–8 on each side in males, separated by 8–11 poreless scales from precloacal pore series; enlarged femoral scales present; postcloacal spurs 2 or 3; subcaudal scales transversely enlarged; lamellae under toe IV 18–22; dorsal pattern consisting of a dark nuchal loop, a continuous or partly interrupted neck band, and five or six in part irregular transverse body bands between limb insertions.


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CommentHabitat: Specimens were found at night between 18:30 and 21:00, on limestone cliffs and in rock crevices, approximately 0.3–1.5 m above the ground, at elevations between 17–28 m a.s.l. The surrounding habitat consisted of secondary karst forest of medium and small hardwoods mixed with shrubs and vines (Fig. 6). Air temperature ranged from 25 to 32.1oC and relative humidity was 70–90%.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyWe name this new species in honor of our colleague and friend, Assoc. Prof. Dr. Son Lan Hung Nguyen, Faculty of Biology, Hanoi National University of Education, Vietnam, in recognition of his support of our research and conservation work in Vietnam. 
References
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • LE, DZUNG TRUNG; TRUONG QUANG NGUYEN, MINH DUC LE, THOMAS ZIEGLER 2016. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Binh Province, Vietnam. Zootaxa 4162 (2): 268–282 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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