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Cyrtodactylus thylacodactylus MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Phnom Dalai, Pocket-fingered, Bent-toed Gecko 
SynonymCyrtodactylus thylacodactylus MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER 2019: 38
Cyrtodactylus intermedius — DALTRY & CHHEANG in DALTRY & MOMBERG 2000: 107
Cyrtodactylus intermedius — NEANG et al. 2010: 137 
DistributionCambodia (Pursat: Samkos Wildlife Sanctuary)

Type locality: Camp 1, Phnom Dalai, Pursat Province, Cambodia (1225’23”N, 10304’07”E, 556 m in elevation).  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 9319, adult male, collected on 6 July 2009 by Neang Thy, L. Lee Grismer, Chan Kin Onn, Jesse L. Grismer, Perry L. Wood Jr., and Timothy M. Youmans. Paratypes. Juvenile male LSUHC 9318 bears the same collection data as the holotype; adult females LSUHC 9326, 9336 and juvenile female LSUHC 9325 were collected on 7 July 2009, by the same collectors at camp 2, Phnom Dalai, Pursat Province, Cambodia (1226’13”N, 10303’21”E, 968 m in elevation). Juvenile female 9349 was collected from camp 2 by the same collectors on 8 July 2009. 
DiagnosisDiagnosis: Adult males reaching 71.5 mm SVL, adult females reaching 74.6 mm SVL; seven supralabials, 8– 10 infralabials; 29–31 paravertebral tubercles; 18–20 longitudinal rows of dorsal tubercles; 36–42 rows of ventral scales; five or six expanded subdigital lamellae proximal to the digital inflection, 10–11 unmodified, distal, subdigital lamellae; 15–18 total subdigital lamellae on fourth toe; enlarged femoral and precloacal continuous; 17– 22 enlarged femoral scales; proximal femoral scales the same size as distal femorals; 7–9 enlarged precloacal scales with pores on each in males; three rows of enlarged post-precloacal scales; 1–3 postcloacal tubercles; deep interdigital pocketing on either side of digit III on forefoot; deep interdigital pocketing present between third and fourth digits of hind feet; dark pigmented blotches absent from top of head; posterior border of nuchal loop rounded; and four or five dark body bands (summarized in Tables 7 + 12 in Murdoch et al. 2019).


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CommentHAbitat: evergreen hill forest in the vicinity of rocky streams. All specimens were collected at night at the base of trees and on leaves and branches 1–1.5 m above the ground. Lizards were most commonly seen at the higher elevations of Camp 11. 
EtymologyThe specific epithet, thylacodactylus, is a Latinized adjective in masculine gender, derived from Greek thylakos meaning “pouch”, and daktylos meaning “digit” in reference to the unique morphology of the new species having deep interdigital pocketing between some of the digits. 
References
  • CHHIN, S., NEANG, T., CHAN, S., KONG, K., OU, R., SAMORN, V., ... & GRISMER, L. L. 2024. A new species in the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from an isolated limestone karst formation in southwestern Cambodia. Zootaxa 5474 (1): 1-20
  • Daltry, J.C. & Momberg, F. (Eds.), 2000. Cardamom Mountains Biodiversity Survey 2000. Fauna & Flora International, Cambridge, pp. 252
  • GRISMER, L. LEE; CHAN KIN ONN, JAMIE R. OAKS, THY NEANG, LANG SOKUN,<br>MATTHEW L. MURDOCH, BRYAN L. STUART & JESSE L. GRISMER 2020. A new insular species of the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from Cambodia with a discussion of habitat preference and ecomorphology. Zootaxa 4830 (1): 075–102 - get paper here
  • MURDOCH, MATTHEW L.; L. LEE GRISMER, PERRY L. JR. WOOD, THY NEANG, NIKOLAY A. POYARKOV, NGO VAN TRI, ROMAN A. NAZAROV, ANCHALEE AOWPHOL, OLIVIER S.G. PAUWELS, HUNG NGOC NGUYEN, JESSE L. GRISMER 2019. Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa 4554 (1): 001–062 - get paper here
  • Neang Thy, L.L. Grismer et al. 2010. First report of the herpetofauna of Dalai Mountain in Phnom Samkos Wildlife Sactuary, southwestern Cardamon Mountains, Cambodia. Cambodian J Nat. Hist. 2010 (2): 127-143 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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