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Cyrtodactylus amphipetraeus CHOMDEJ, SUWANNAPOOM, PAWANGKHANANT, PRADIT, NAZAROV, GRISMER & POYARKOV, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Tak bent-toed Gecko
Thai: Tuk kai tak 
SynonymCyrtodactylus amphipetraeus CHOMDEJ, SUWANNAPOOM, PAWANGKHANANT, PRADIT, NAZAROV, GRISMER & POYARKOV 2020 
DistributionThailand (Tak)

Type locality: limestone rocks at the entrance to the Tham Sri Fah Cave, Mae Sot District, Tak Province, Thailand (16.602162°N, 98.712481°E WGS; 710 m in elevation  
Reproduction 
TypesHolotype. AUP 00696 (Agriculture and Natural Resources, University of Phayao (Phayao, Thailand), adult male, collected on August 3, 2019, at 21:00 hrs by S. Chomdej, C. Suwannapoom and P. Pawangkhanant.
Paratypes. Nine specimens, including the paratypes AUP-00688–90 (three adult males), and AUP-00691–93 and AUP-0097 (four adult females) bear the same collecting data as the holotype; AUP-00698, an adult male, was collected on August 3, 2019, at 20:00 hrs by S. Chomdej, C. Suwannapoom and P. Pawangkhanant from granite rocks near the Tha Ra Rak waterfall, Mae Sot District, Tak Province, Thailand (16.569339° N, 98.694566° E WGS; 610 m in elevation); ZMMU R-16626 (field No. NAP-06637) was collected on November 13, 2016, at 20:00 hrs by N.A. Poyarkov and P. Pawangkhanant from granite rocks near the Tha Ra Rak waterfall, Mae Sot District, Tak Province, Thailand (16.569339° N, 98.694566° E WGS; 610 m in elevation). 
DiagnosisDiagnosis: Cyrtodactylus amphipetraeus sp. nov. differs from all species in the C. sinyineensis group by having the combination of nine supralabials; seven infralabials; 34–38 paravertebral tubercles; 17–20 longitudinal rows of dorsal tubercles; 28–30 ventral scales ventral scales; seven expanded subdigital lamellae on the fourth toe; 11 or 12 unmodified subdigital lamellae on the fourth toe; 18 or 19 total subdigital lamellae on the fourth toe; 27–34 enlarged femoral scales; a total of 10–12 pore-bearing femoral scales in males; 8–11 enlarged precloacal scales; 7–9 pore-bearing precloacal scales in males; three rows of enlarged post-precloacal scales; approximately 4–7 broken to hour glass shaped dorsal body bands; 10–12 light-colored caudal bands (n=2); 11–13 dark-colored caudal bands (n=2); raised and strongly keeled dorsal tubercles that extend beyond base of tail; enlarged femoral and precloacal scales nearly the same size and continuous; pore-bearing femoral and precloacal scales not continuous; medial subcaudals two to three times wider than long and extending onto lateral side of tail; iris green; nuchal loop lacking an anterior azygous notch, and bearing a jagged posterior border; dorsal bands bearing paravertebral elements, generally equal in width than interspaces, bearing lightened centers, edged with white tubercles; dark markings in dorsal interspaces; light caudal bands in adults bearing dark-colored markings; light-colored caudal bands not encircling tail; and mature regenerated tail not spotted (Table 5 in CHOMDEJ et al. 2020).


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Comment 
EtymologyThe specific epithet amphipetraeus is a Latinized adjective in nominative singular, derived from Greek amphi or ἀμφί (meaning of both kinds) and petra or πέτρα (for rock). The species name is given in reference to the remarkable natural history of this species which inhabits both limestone and granite rocks. 
References
  • CHOMDEJ, S., SUWANNAPOOM, C., PAWANGKHANANT, P., PRADIT, W., NAZAROV, R. A., GRISMER, L. L., & POYARKOV, N. A. 2020. A new species Cyrtodactytlus Gray (Squamata: Gekkonidae) from western Thailand and the phylogenetic placement of C. inthanon and C. doisuthep. Zootaxa 4838 (2): 179-209 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
 
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