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Cyrtodactylus gubaot WELTON, SILER, LINKEM, DIESMOS & BROWN, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus gubaot WELTON, SILER, LINKEM, DIESMOS & BROWN 2010
Cyrtodactylus agusanensis — BROWN & ALCALA 1978: 16 (part) 
DistributionPhilippines (Leyte)

Type locality: Leyte Island, Leyte Province, Municipality of Baybay, Barangay Pilim, Sitio San Vicente (10.731° N, 124.826° E, 82 m elevation; WGS84.  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: PNM 9724 (formerly KU 311184; field no. CDS 3273), adult male, collected on 3 November 2007 by CDS and J. Fernandez. 
DiagnosisDiagnosis: Cyrtodactylus gubaot most closely resembles congeners from the C. agusanensis complex and is distinguished from C. agusanensis by midbody dorsals 92–105 (vs. 111–124), paravertebrals 150–162 (vs. 184–196), postcloacal lateral tubercles 4–6 (vs. 8–11), venter dark gray with white flecks (vs. cream throughout), the presence (vs. absence) of a canthal stripe, anterior–posterior dorsal band projections moderate (vs. extensive), and cephalic tubercles large (vs. moderate).
Cyrtodactylus gubaot is diagnosed from sumuroi (Samar) by SVL 85.6–100.1 mm (vs. 73.9–84.9), paravertebrals 150–162 (vs.163–180), venter dark gray with white flecks (vs. cream throughout), anterior–posterior dorsal band projections moderate (vs. minimal), dark dorsal bands not enclosing light bands (vs. light bands enclosed), cephalic tubercles large (vs. moderate), and caudal annuli with dorsal tubercles 9–12 (vs. 3–7). Cyrtodactylus gubaot also exhibits tendencies toward a greater number of midbody ventrals (54–67 vs. 53– 58), midbody dorsals (92–105 vs. 87–98), and subdigital lamellae beneath Finger III (20–26 vs. 18–22) and Toe IV (24–30 vs. 21–24).
Cyrtodactylus gubaot is readily diagnosed from C. mamanwa (Dinagat) by paraverte- brals 150–162 (vs. 173–192), venter dark gray with white flecks (vs. cream throughout), the presence of a canthal stripe (vs. absence), dark dorsal bands not enclosing light bands (vs. light bands enclosed), and cephalic tubercles large (vs. small). Cyrtodactylus gubaot also shows a tendency toward fewer paravertebral tubercles (19–27 vs. 25–31).
Cyrtodactylus gubaot is distinguished from all other Philippine congeners by characters of body size and scalation (Table 3). Cyrtodac- tylus gubaot can be distinguished from C. annulatus, C. tautbatorum, C. jambangan, C. redimiculus, and C. philippinicus by dark gray venter with white flecks, larger cephalic tubercle size, and the presence (vs. absence) of a canthal stripe; from C. annulatus, C. tautbatorum, C. jambangan, and C. redimicu- lus by a greater number of subdigital lamellae beneath Toe IV; from C. annulatus, C. tautbatorum, C. jambangan, and C. philippi- nicus by the presence of femoral pore-bearing scales; from C. annulatus, C. tautbatorum, and C. redimiculus by a greater number of midbody tubercle rows; from C. redimiculus and C. philippinicus by dark dorsal bands enclosing light bands (vs. light bands not enclosed); from C. tautbatorum and C. jambangan by larger SVL; from C. annulatus and C. tautbatorum by more postcloacal lateral tubercles; from C. tautbatorum by a greater number of subdigital lamellae beneath Finger III; from C. jambangan by a greater number of caudal annuli with dorsal tubercles; and from C. redimiculus by a greater number of midbody ventrals, fewer paravertebrals, and moderate anterior–posterior dorsal band pro- jections (vs. minimal). 
Comment 
EtymologyThe specific epithet is a combination of the Leyte dialect (Waray-waray) terms guba (forest) and buot (spirit, essence) in reference to our observation that the new species is one of the most abundant and commonly observed reptiles from the forests of central Leyte Island. 
References
  • Brown W C; Alcala A C 1978. Philippine lizards of the family Gekkonidae. Silliman University, Dumaguete City, Philippines 1978: i-vii, 1-146
  • Decena, S. C. P., Macasait Jr, D. R., & Arguelles, M. S. 2023. Species Accounts, Assemblage, and Microhabitats of Amphibians and Reptiles of Northeastern Leyte, Philippines. Philippine Journal of Science, 152(1), 1-34
  • Oaks, J. R., Siler, C. D. and Brown, R. M. 2019. The comparative biogeography of Philippine geckos challenges predictions from a paradigm of climate‐driven vicariant diversification across an island archipelago. Evolution doi:10.1111/evo.13754 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Charles W. Linkem, Arvin C. Diesmos, and Rafe M. Brown 2010. Philippine Bent-Toed Geckos of the Cyrtodactylus agusanensis Complex: Multilocus Phylogeny, Morphological Diversity, and Descriptions of Three New Species. Herpetological Monographs 24 (1): 55-85 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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