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Cyrtodactylus intermedius (SMITH, 1917)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Intermediate Bow-fingered Gecko, Cardamon Forest Gecko
G: Kleiner Bogenfingergecko 
SynonymGymnodactylus intermedius SMITH 1917: 221
Cyrtodactylus intermedius — TAYLOR 1963: 734
Gymnodactylus (Cyrtodactylus) intermedius — WERMUTH 1965: 54
Cyrtodactylus intermedius — COX et al. 1998: 86
Cyrtodactylus (Cyrtodactylus) intermedius — RÖSLER 2000: 66
Cyrtodactylus intermedius — DEWAND 2005: 140
Cyrtodactylus intermedius — NGUYEN et al. 2009
Cyrtodactylus intermedius — MURDOCH et al. 2019 
DistributionSE Thailand (Province Chanthaburi, Khao Yai), Cambodia, Koh Chang, Vietnam

Type locality: Khao Sebab, Chantabun, Southeast-Siam.  
Reproductionoviparous 
TypesSyntype: UMMZ 78687, MCZ R-39040, collected by M.A. Smith, March 1916 
DiagnosisDiagnosis: Adult males reaching 78.7 mm SVL; eight supralabials, 9 or 10 infralabials; 31–33 paravertebral tubercles; 19 or 20 longitudinal rows of dorsal tubercles; 42–44 rows of ventral scales; seven or eight expanded subdigital lamellae proximal to the digital inflection, 12–14 unmodified, distal, subdigital lamellae; 19–21 total subdigital lamellae on fourth toe; enlarged femoral and precloacal scales continuous; 23 or 24 enlarged femoral scales; proximal femoral scales one-half the size of distal femorals; 8–10 enlarged precloacal scales with pores on each in males; three rows of enlarged post-precloacal scales; two or three postcloacal tubercles; no pocketing between digits of hind or forefeet; dark pigmented blotches absent on top of head; posterior border of nuchal loop rounded; and four dark body bands (Tables 7-8 in Murdoch et al. 2019). 
CommentGroup: Grismer et al. 2023: 123 (Table 5) provide summary statistics of meristic traits for the members of the intermedius group, consisting of disjunctus, hontreensis, kohrongensis, auralensis, bokorensis, cardamomensis, thylacodactylus, laangensis, septimontium, phuquocensis, and intermedius. 
References
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Bulian, Joachim 2013. Reptiliensuche mit Hindernissen - Teil 2. Terraria-Elaphe 2013 (3): 44-45 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • CHHIN, S., NEANG, T., CHAN, S., KONG, K., OU, R., SAMORN, V., ... & GRISMER, L. L. 2024. A new species in the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from an isolated limestone karst formation in southwestern Cambodia. Zootaxa 5474 (1): 1-20
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • David, P.; Teynié, A. & Ohler, A. 2004. A new species of Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae) from southern Laos. Raffles Bull. Zool. 52 (2): 621-627 - get paper here
  • DAVID, PATRICK; TRUONG QUANG NGUYEN, NICOLE SCHNEIDER & THOMAS ZIEGLER 2011. A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa 2833: 29–40 - get paper here
  • Derwanz, K. 2005. Thailand – Dschungelabendteuer Khao Yai. Opferkuch GmbH, Aalen/Ellwangen, 269 pp
  • Geissler, P.; Nazarov, R.; Orlov, N.L.; Böhme, W.; My Phung, T.; Nguyen, T.Q. & Ziegler, T. 2009. A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 2161: 20–32 - get paper here
  • Goldberg, Stephen R. 2012. Notes on reproduction of the Cardamon forest gecko, Cyrtodactylus intermedius (Squamata: Gekkonidae) from Cambodia and Thailand. Hamadryad 36 (1): 58-60 - get paper here
  • GRISMER, L. L., PAWANGKHANANT, P., IDIIATULLINA, S. S., TROFIMETS, A. V., NAZAROV, R. A., SUWANNAPOOM, C., & POYARKOV, N. A. 2023. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the Thai-Malay Peninsula and the independent evolution of cave ecomorphology on opposite sides of the Gulf of Thailand. Zootaxa 5352(1): 109-136 - get paper here
  • GRISMER, L. LEE; CHAN KIN ONN, JAMIE R. OAKS, THY NEANG, LANG SOKUN,<br>MATTHEW L. MURDOCH, BRYAN L. STUART & JESSE L. GRISMER 2020. A new insular species of the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from Cambodia with a discussion of habitat preference and ecomorphology. Zootaxa 4830 (1): 075–102 - get paper here
  • Grismer, L. Lee; Thy Neang, Thou Chav, Perry L. Wood, Jr., Jamie R. Oaks, Jeremy Holden, Jesse L. Grismer, Thomas R. Szutz and Timothy M. Youmans 2008. Additional amphibians and reptiles from the Phnom Samkos Wildlife Sanctuary in Northwestern Cardamom Mountains, Cambodia, with comments on their taxonomy and the discovery of three new species. Raffles Bulletin of Zoology 56 (1): 161-175 - get paper here
  • Grismer, L.L. et al. 2007. THE HERPETOFAUNA OF THE PHNOM AURAL WILDLIFE SANCTUARY AND CHECKLIST OF THE HERPETOFAUNA OF THE CARDAMOM MOUNTAINS, CAMBODIA. Hamadryad 31 (2): 216 – 241 - get paper here
  • Grismer, L.L., Neang, T., Chav, T. & Grismer, J.L. 2008. Checklist of the amphibians and reptiles of the Cardamom region of Southwestern Cambodia. Cambodian Journal of Natural History 2008(1): 12–28 - get paper here
  • Krysko, K. L.; Rehman, H. & Auffenberg, K. 2007. A new species of Cyrtopodion (Gekkonidae: Gekkoninae) from Pakistan. Herpetologica 63 (1): 100-113 - get paper here
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickford, and Rafe M. Brown 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; TRUONG QUANG NGUYEN, HUY QUANG DO, THOMAS ZIEGLER, 2011. A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa 3129: 39–50 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • LUU, VINH QUANG;THOMAS CALAME, MICHAEL BONKOWSKI, TRUONG QUANG NGUYEN & THOMAS ZIEGLER 2014. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa 3760 (1): 054–066 - get paper here
  • Nabhitabhata, J., T. Chan-ard and Y. Chuaynkern 2000. Checklist of Amphibians and Reptiles in Thailand. Office of Environmental Policy and Planning, Bankok. 152pp.
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  • NAZAROV, ROMAN; NIKOLAY A. POYARKOV, NIKOLAI L. ORLOV, TRUNG MY PHUNG, TAO THIEN NGUYEN, DUC MINH HOANG & THOMAS ZIEGLER 2012. Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 3302: 1–24 - get paper here
  • Neang Thy, L.L. Grismer et al. 2010. First report of the herpetofauna of Dalai Mountain in Phnom Samkos Wildlife Sactuary, southwestern Cardamon Mountains, Cambodia. Cambodian J Nat. Hist. 2010 (2): 127-143 - get paper here
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  • Schäfer, C. 1993. Cyrtodactylus intermedius (SMITH, 1917). Sauria 15 Suppl.: 279–282 - get paper here
  • Smith, M.A. 1917. Descriptions of new reptiles and a new batrachian from Siam. J. Nat. Hist. Soc. Siam 2: 221-225 - get paper here
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  • Tri, Ngo Van & Onn, Chan Kinn 2010. A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, Southern Vietnam. Zootaxa 2504: 47–60 - get paper here
  • Youmans, Timothy M. & L. Lee Grismer 2006. A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 10 (1): 61-70
  • Ziegler, T.; ROMAN NAZAROV, NIKOLAI ORLOV, TRUONG QUANG NGUYEN, THANH NGOC VU, KIEN NGOC DANG, TRI HUY DINH, ANDREAS SCHMITZ 2010. A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, Truong Son Range, Vietnam. Zootaxa 2413: 20–36 - get paper here
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