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Cyrtodactylus lateralis (WERNER, 1896)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Werner’s Prehensile-tailed Bent-toed Gecko, Sumatra Bow-fingered Gecko, Spiny Forest Gecko
G: Seitenstachel-Bogenfingergecko 
SynonymGymnodactylus lateralis WERNER 1896: 11
Gymnodactylus lateralis — DE ROOIJ 1915: 7
Cyrtodactylus lateralis — MANTHEY & GROSSMANN 1997: 223
Cyrtodactylus (Cyrtodactylus) lateralis — RÖSLER 2000: 66
Cyrtodactylus lateralis — TEYNIÉ et al. 2010 
DistributionWest Malaysia, Indonesia (Sumatra)

Type locality: Sumatra  
Reproductionoviparous 
TypesHolotype: ZMB 12029 
DiagnosisDiagnosis: A large species of Cyrtodactylus reaching 100 mm SVL (LSUHC 12579) and distinguished from all congeners by the following combination of characters: (1) body robust; limbs and digits moderate in length; (2) tail prehensile; intact tails 0.97–1.22 times longer than body; (3) tubercles extending from frontal region and supraorbital skin to cover most of tail, 16–20 irregular longitudinal rows of tubercles at midbody, 21–28 irregular transverse rows between limbs; (4) dorsal antebrachium and brachium tuberculate; (5) tubercles on tail extending to 90% of its length; (6) 51–66 ventrals in a transverse row at midbody; (7) conical, spinose tubercles in ventrolateral fold; (8) subcaudals not transversely enlarged; (9) moderate longitudinal sulcus in precloacal region of males; (10) 9–13 precloacal pores in males (0–15) pore primordia in females), femoral pores absent, precloacal pores sunken into precloacal sulcus; pore secretions red; (11) greatly enlarged precloacal pore-bearing scales present; (12) ventral surface of thighs entirely granular or granular except for 2–13 (count combined for both sides) widely spaced enlarged femoral scales; (13) 18–24 lamellae under fourth toe; (14) cloacal tubercles 1–2 on each side, usually in contact with one another; (15) dorsum gray to brown with dark brown markings; venter pale pink, immaculate or with diffuse, darkly pigmented scales forming no obvious pattern; (16) labials pale, contrasting with darker facial band; occiput with few large blotches; postocular stripe brown, edged dorsally by thin black line then by pale pigmented scales; (17) 5–6 transverse, usually broken bands on body; 7–9 bands on tail; caudal bands complete ventrally though faint (Harvey et al. 2016).


Additional details (2008 characters) are available for collaborators and contributors. Please contact us for details. 
CommentDistribution: See map in Harvey et al. 2016: 526 (Fig. 5). 
References
  • Grismer L. Lee 2005. New species of Bent-toed Gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology 39 (3):424-432. - get paper here
  • Grismer L. Lee & Leong, T.M. 2005. New Species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Peninsular Malaysia. Journal of Herpetology 39 (4): 584-591 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.L.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L.; Wood, P.L. & Youmans, T.M. 2007. Redescription of the gekkonid lizard Cyrtodactylus sworderi (SMITH 1925) from Southern Peninsular Malaysia. Hamadryad 31 (2): 250-257 - get paper here
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • HARVEY, MICHAEL B.; KYLE A. O’CONNELL, ELIJAH WOSTL, AWAL RIYANTO, NIA KURNIAWAN, ERIC N. SMITH, L. LEE GRISMER 2016. Redescription Cyrtodactylus lateralis (Werner) (Squamata: Gekkonidae) and Phylogeny of the Prehensile-tailed Cyrtodactylus. Zootaxa 4107 (4): 517-540 - get paper here
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickford, and Rafe M. Brown 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • Onn, Chan Kin & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • O’Connell, K. A., Smart, U., Sidik, I., Riyanto, A., Kurniawan, N., & Smith, E. N. 2019. Diversification of bent-toed geckos (Cyrtodactylus) on Sumatra and west Java. Molecular Phylogenetics and Evolution, 134, 1–11 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H. & Glaw, F. 2008. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa 1729: 8–22 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Welton, L.J, Siler, C.D., Diesmos, A.C. & Brown, R.M. 2010. Phylogeny-based species delimitation of southern Philippines bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago. Zootaxa 2390: 49–68 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Arvin Diesmos, and Rafe M. Brown 2009. A NEW BENT-TOED GECKO (GENUS CYRTODACTYLUS) FROM SOUTHERN PALAWAN ISLAND, PHILIPPINES AND CLARIFICATION OF THE TAXONOMIC STATUS OF C. ANNULATUS. Herpetologica 65 (3): 328-343 - get paper here
  • Werner, F. 1896. Zweiter Beitrag zur Herpetologie der indo-orientalischen Region. Verh. Zool. Bot. Ges. Wien 46: 6-24 - get paper here
  • Youmans, Timothy M. & L. Lee Grismer 2006. A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 10 (1): 61-70
 
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